Jain, V., Bugnyar, Th., Cunningham, S.J., Gallego Abenza, M., Loretto, M.-C., Sumasgutner, P.(2022). The spatial and temporal exploitation of anthropogenic food sources by common ravens (Corvus corax) in the Alps. Movement Ecology.

Horn, L., Cimarelli, G., Boucherie, P.H., Šlipogor, V. and Bugnyar, T., 2022. Beyond the dichotomy between field and lab—the importance of studying cognition in context. Current Opinion in Behavioral Sciences46, p.101172.

Boucherie, P.H., Gallego-Abenza, M., Massen, J.J. and Bugnyar, T., 2022. Dominance in a socially dynamic setting: hierarchical structure and conflict dynamics in ravens' foraging groups. Philosophical Transactions of the Royal Society B377(1845), p.20200446.

Miller, R., Lambert, M.L., Frohnwieser, A., Brecht, K.F., Bugnyar, T., Crampton, I., Garcia-Pelegrin, E., Gould, K., Greggor, A.L., Izawa, E.I. and Kelly, D.M., 2022. Socio-ecological correlates of neophobia in corvids. Current Biology32(1), pp.74-85.

Gallego-Abenza, M., Boucherie, P.H. and Bugnyar, T., 2022. Early social environment affects attention to social cues in juvenile common ravens, Corvus corax. Royal Society Open Science9(6), p.220132.

Gallup, A.C., Schild, A.B., Ühlein, M.A., Bugnyar, T. and Massen, J.J., 2022. No Evidence for Contagious Yawning in Juvenile Ravens (Corvus corax): An Observational Study. Animals12(11), p.1357.

Puehringer-Sturmayr, V., Krejci, J., Schuster, R., Kleindorfer, S., Kotrschal, K., Frigerio, D., & Loretto, M. (2022). Space use and site fidelity in the endangered Northern Bald Ibis Geronticus eremita: Effects of age, season, and sex. Bird Conservation International, 1-13.

Stofberg, M., Amar, A., Sumasgutner, P., Cunningham, S.J. (2022). Staying cool and eating junk: Influence of heat dissipation and anthropogenic food on foraging and body condition in an urban passerine. Landscape and Urban Planning 226. The article can be accessed for 50 days

Kleindorfer, S., Colombelli-Négrel, D., Common, L.K., O'Connor, J.A., Peters, K.J., Katsis, A.C., Dudaniec, R.Y., Sulloway, F.J., Adreani, N.M. (2022). Functional traits and foraging behaviour: Avian vampire fly larvae change the beak and fitness of their Darwin's finch hosts. Functional Ecology, 00, 1–12.

Adreani, N.M., Valcu, M., Citizen Scientists, Mentesana, L. (2022). Asymmetric architecture is non-random and repeatable in a bird's nest. Current Biology 32(9): 412-413.

García-Loor J (2022) Elevational range extension of Drab Water Tyrant, Ochthornis littoralis (Pelzeln, 1868) (Passeriformes, Tyrannidae). Check List 18(3): 451-453.

Gegendorfer, G., Rittenschober, J., Vesely, H. and Frigerio D. (2022). Volunteering In Monitoring Bird Behavior: Accuracy Matters! Front.Young Minds. doi: 10.3389/frym.2022.596181

Naikatini, A.N., Keppel, G., Brodie G, Kleindorfer, S. (2022). Avian diversity and abundance across years: consistent patterns in forests but not grasslands on Viti Levu, Fiji. Pacific Conservation Biology.

Naikatini, A.N., Keppel, G., Brodie, G., Kleindorfer, S., 2022. Interspecific Competition and Vertical Niche Partitioning in Fiji’s Forest Birds. Diversity.

Guggenberger, M., Adreani, N.M., Foerster, K., Kleindorfer, S., 2022. Vocal recognition of distance calls in a group-living basal bird: the greylag goose, Anser anser. Animal behaviour.


Miller, R., Lambert, M.L., Frohnwieser, A., Brecht, K.F., Bugnyar, T., Crampton, I., Garcia-Pelegrin, E., Gould, K., Greggor, A.L., Izawa, E.I. and Kelly, D.M., 2021. Socio-ecological correlates of neophobia in corvids. Current Biology.
Ersoy, S., Maag, N., Boehly, T., Boucherie, P.H. and Bugnyar, T., 2021. Sex-specific parental care during postfledging in common ravens. Animal Behaviour, 181, pp.95-103.
Stocker, M., Prosl, J., Vanhooland, L.C., Horn, L., Bugnyar, T., Canoine, V. and Massen, J.J., 2021. Measuring salivary mesotocin in birds-Seasonal differences in ravens' peripheral mesotocin levels. Hormones and Behavior, 134, p.105015.
Wenig, K., Boucherie, P.H. and Bugnyar, T., 2021. Early evidence for emotional play contagion in juvenile ravens. Animal Cognition, pp.1-13.
Boucherie, P.H., Cimarelli, G., Horn, L., Šlipogor, V. and Bugnyar, T., 2021. Cognition in Context. In Proceedings of the Annual Meeting of the Cognitive Science Society (Vol. 43, No. 43).

Katsis AC, Colombelli-Négrel D, Common LK, O’Connor JA, Dudaniec RY, Garcia-Loor J, Kleindorfer S (2021). Nestling behaviour predicts naris deformation in Darwin’s finches parasitised by the avian vampire fly. Biological Journal of the Linnaean Society blab092.

Katsis A, Buchanan K, Kleindorfer S, Marriette M (2021). Long-term effects of prenatal sound experience on songbird behavior and their relation to song learning. Behavioral Ecology and Sociobiology 75(1), 1-13.

Antonson ND, Rivera M, Abolins-Abols M, Kleindorfer S, Liu W-C, Hauber ME (2021). Early auditory experience alters genome-wide methylation in the auditory forebrain of songbird embryos. Neuroscience Letters 755: 135917.

Colombelli-Négrel D, Hauber ME, Evans C, Katsis AC, Brouwer L, Adreani NM, Kleindorfer S (2021). Prenatal auditory learning in avian vocal learners and non-learners. Philosophical Transactions of the Royal Society 376: 20200247.

Common LK, Sumasgutner P, Sumasgutner S, Colombelli-Négrel D, Dudaniec RY, Kleindorfer S (2021). Temporal and spatial variation in sex-specific abundance of the avian vampire fly (Philornis downsi). Parasitology Research

Wascher, C.A., Allen, K. and Szipl, G., 2021. Learning and motor inhibitory control in crows and domestic chickens. Royal Society Open Science, 8(10), p.210504.

Wascher, C. A. F., Baur, N., Hengl, M., Köck, C., Pegger, T., Schindlbauer, J., & Wemer, L. (2021). Behavioral responses of captive corvids to the presence of visitors. Animal Behavior and Cognition, 8(4), 481-492.

Kleindorfer S, Common LK, O’Connor JA, Garcia-Loor J, Katsis AC, Dudaniec RY, Colombelli-Négrel D, Adreani NM (in press, will be published 15 Dec). Female in-nest attendance predicts the number of ectoparasites in Darwin’s finch species. Proceedings of the Royal Society B.

Odom, K.J., Cain, K.E., Hall, M.L., Langmore, N.E., Mulder, R.A., Kleindorfer, S., Karubian, J., Brouwer, L., Enbody, E.D., Jones, J.A. and Dowling, J.L. (2021). Sex role similarity and sexual selection predict male and female song elaboration and dimorphism in fairy‐wrens. Ecology and Evolution.

Kleindorfer, S., Common, L.K., Sumasgutner, P. (2021). Nesting Success and Nesting Height in the Critically Endangered Medium Tree Finch (Camarhynchus pauper). Birds MDPI.

Nebel C*, Sumasgutner P, Rodseth E, Ingle R, Childs DZ, Curtis-Scott O, Amar A (2021): Multigenerational pedigree of wild individually marked black sparrowhawks suggests that dark plumage colouration is dominantly inherited. Journal of Zoology.

Sumasgutner P (2021): Editorial: A Southern perspective on urban birds. Ostrich: Journal of African Ornithology 92: III–V.

Common LK, Sumasgutner P, Dudaniec RY, Colombelli-Négrel D, Kleindorfer S*: Avian vampire fly (Philornis downsi) mortality differs across Darwin’s finch host species. Scientific Reports 11:15832.

Frigerio D, Sumasgutner P, Kotrschal K, Kleindorfer S*, Hemetsberger S (2021): From individual to population level: Temperature and snow cover modulates fledging success through breeding phenology in Greylag geese (Anser anser). Scientific Reports 11:16100.

Catto S, Sumasgutner P*, Amar A, Thomson R, Cunningham S (2021): Pulses of anthropogenic food availability appear to benefit parents, but compromise nestling growth in urban Red-winged Starlings. Oecologia

Wemer L, Hegemann A, Isaksson C, Nebel Carina, Kleindorfer S, Gamauf A, Adrion M, Sumasgutner P* (2021): Reduced ectoparasite load, body mass and blood haemolysis in Eurasian kestrels (Falco tinnunculus) along an urban-rural gradient. The Science of Nature/Naturwissenschaften

Bates, A. E., Primack, R. B., PAN-Environment Working Group, Duarte C. M. (2021) Global COVID-19 lockdown highlights humans as both threats and custodians of the environment. Biological Conservation, 109175

Nebel, C., Amar, A., Hegemann, A., Isaksson C., Sumasgutner, P. (2021) Parental morph combination does not influence innate immune function in nestlings of a colour-polymorphic African raptor. Scientific Reports, 11: 11053

Sumasgutner, P., Buij, R., McClure, C. J. W., Shaw, P., Dykstra, C. R., Kumar, N., Rutz, C. (2021) Raptor research during the COVID-19 pandemic provides invaluable opportunities for conservation biology. Biological Conservation, 109149

McPherson, S. C., Sumasgutner, P., Downs, C. T. (2021) South African raptors in urban landscape: a review. Journal of African Ornithology, 92(1): 41-57

Reynolds, C., Byrne, M. J., Chamberlain, D. E., Howes, C. G., Seymour, C. L., Sumasgutner, P., Taylor, P. J. (2021) Urban Animal Diversity in the Global South. In: Shackleton, C. M., Cilliers, S. S., Davoren, E., du Toit, M. J. (Eds) Urban Ecology in the Global South. Switzerland: Springer International Publishing. pp 169-202

Westreicher, F., Cieslinski, M., Ernst, M., Frigerio, D., Heinisch, B., Hübner, T., Rüdisser, J. (2021) Recognizing failures in citizen science projects: Lessons learned. 6th Austrian Citizen Science Conference 2020, Proceedings of Science (ACSC2020), 393

Neuböck-Hubinger, B., Grubner, K., Frigerio, D., Hirschenhauser, K. (2021) Natural science education - a claim for equality, diversity and inclusion in Citizen Science activities. 6th Austrian Citizen Science Conference 2020, Proceedings of Science (ACSC2020), 393

Rittenschober, J., Puehringer-Sturmayr, V., Frigerio, D. (2021) The relevance of a digital platform for the citizen science project NestCams. 6th Austrian Citizen Science Conference 2020, Proceedings of Science (ACSC2020), 393

Risi, T. C., Sumasgutner, P., Cunningam, S. J. (2021) Anthropogenic food availability and body mass maintenance in urban Red-winged Starlings Onychognathus morio. Ostrich, 1-10

Colombelli-Négrel, D., Kleindorfer, S. (2021) Behavioural response to songs between genetically diverged allopatric populations of Darwin's small tree finch in the Galápagos. Journal of Evolutionary Biology, 00: 1-14

Mentesana, L., Adreani, N. M. (2021) Acute aggressive behavior perturbates the oxidative status of a wild bird independently of testosterone and progesterone. Hormones and Behavior, 128: 104913

Katsis, A., Buchanan, K., Kleindorfer, S., Marriette, M. (accepted) Long-term effects of prenatal sound experience on songbird behavior and their relation to song learning. Behavioral Ecology and Sociobiology

Frigerio, D., Richter, A., Per, E., Pruse, B., Vohland, K. (2021) Citizen Science in natural sciences. In: Vohland, K., Land, A., Ceccaroni, L., Lemmens, R., Perelló, J., Ponti, M., Samson, R., Wagenknecht, K. (eds) The Science of Citizen Science. Springer, Chapter 5,

Gallego-Abenza, M., Blum, C., Bugnyar, T. (2021) Who is crying wolf? Seasonal effect on anti-predator response to age-specific alarm-calls in common ravens, Corvus corax. Learning & Behavior. doi:



Davidkova, M., Vesely, P., Syrova, M., Nacarova, J., Bugnyar, T. (2020) Raven respond to unfamiliar corvid alarm calls. Journal of Ornithology, 161: 967-975

Beck, K., Loretto, M.-C., Bugnyar, T. (2020) Effects of site fidelity, group size and age on food caching behaviour of common ravens, Corvus corax. Animal Behaviour, 164: 51-64

Loretto, M.-C., Schuster, R., Federspiel, I. G., Heinrich, B., Bugnyar, T. (2020) Contextual imitation in juvenile common ravens (Corvus corax). Animal Behaviour, 163: 127-134

Sierro, J., Loretto, M.-C., Szipl, G., Massen, J. J. M., Bugnyar, T. (2020) Food calling in ravens (Corvus corax) revisited: who is addressed? Ethology, 126(2): 257-266

Kleindorfer, S., Dudaniec, R. Y. (2020) Hybridization fluctuates with rainfall in Darwin's tree finches. Biological Journal of the Linnaean Society, 130(1): 79-88. doi:

Adreani, N. M., D'Amelio, P. B., Gahr, M., ter Maat, A. (2020) Life-Stage Dependent Plasticity in the Auditory System of a Songbird is Signal and Emitter-Specific. Frontiers in Neuroscience, 14:588672. doi: 10.3389/fnins.2020.588672

Böhm, C., Bowden, C. G. R., Seddon, P. J., Hatipoglu, T., Oubrou, W., El Bekkay, M., Quevedo, M. A., Fritz, J., Yeniyurt, C., Lopez, J. M., Orueta, J. F., Frigerio, D., Unsöld, M. (2020) The northern bald ibis Geronticus eremita: history, current status and future perspectives. Oryx, 1-13. doi: 10.1017/S0030605320000198

Heigl, F., Kieslinger, B., Paul, K. T., Uhlik, J., Frigerio, D., Dörler, D. (2020) Co-Creating and Implementing Quality Criteria for Citizen Science. Citizen Science: Theory and Practice, 5(1): p.23.

Gegendorfer, G., Frigerio, D. (2020) How having a family improves digestion in social geese. Frontiers for Young Minds, 8: 540008. doi: 10.3389/frym.2020.540008

Puehringer-Sturmayr, V., Stiefel, T., Kotrschal, K., Kleindorfer, S., Frigerio, D. (2020) Social interactions change with season and age in Northern Bald Ibis. Journal of Ornithology, 162:277–288.

McCarren, S., Sumasgutner, P., Tate, G., Koeslag, A., & Amar, A. (2020). Clinal variation in the polymorphic Black Sparrowhawk Accipiter melanoleucus is unrelated to infection by the blood parasite Haemoproteus nisi. Journal of Ornithology.

Puehringer-Sturmayr, V., Hemetsberger, F., Frigerio, D. (2020) Tracking Birds: How a Little Backpack Could Affect the Life of a Wild Bird. Froniers for Young Minds, 8:116

Mentesana, L., Moiron, M., Guedes, E., Cavalli, E., Tassino, B., Adreani, N. M. (2020) Defending as a unit: sex- and context-specific territorial defence in a duetting bird. Behavioral Ecology and Sociobiology, 74:111

Huchler, K., Schulze, C. H., Gamauf, A., Sumasgutner, P. (2020) Shifting Breeding Phenology in Eurasian Kestrels Falco tinnunculus: Effects of Weather and Urbanization. Frontiers in Ecology and Evolution, 8: 247

Nebel, C., Sumasgutner, P., McPherson, S. C., Tate, G. J., Amar, A. (2020) Contrasting parental color morphs increase regularity of prey deliveries in an African raptor. Behavioral Ecology, 31:1142–1149. Doi: 10.1093/beheco/araa063

Sumasgutner, P., Jenkins, A., Amar, A., Altwegg, R. (2020) Nest boxes buffer the effects of climate on breeding performance in an African urban raptor. PLoS ONE, 15: e0234503

Mueller, R., Amar, A., Sumasgutner, P., McPherson, S. C., Downs, C. (2020) Contrasting effects of urbanization on breeding performance measures in the threatened African Crowned Eagle (Stephanoaetus coronatus). The Condor, duaa024

Frigerio, D., Puehringer-Sturmayr, V., Steinbacher, C., Rittenschober, J., Kleindorfer, S. (2020). Citizen science, animal behaviour and digital devices: challenges and solutions. 5th Austrian Citizen Science Conference, 2019 June 26-28, Obergurgl, Austria. PoS(ACSC2019)002

Common, L.K., O’Connor, J.A., Dudaniec, R.Y., Peters, K.J., Kleindorfer, S. (2020). Evidence for rapid downward fecundity selection in an ectoparasite (Philornis downsi) with earlier host mortality in Darwin’s finches. Journal of Evolutionary Biology.

Puehringer-Sturmayr, V., Loretto, M.C., Hemetsberger, J., Czerny, T., Gschwandegger, J., Leitsberger, M., Kotrschal, K., Frigerio, D. (2020). Effects of bio-loggers on behaviour and corticosterone metabolites of Northern Bald Ibises (Geronticus eremita) in the field and in captivity. Anim Biotelemetry 8: 2. doi:10.1186/s40317-019-0191-5



Nebel, C., Harl, J., Pajot, A., Weissenböck, H., Amar, A., & Sumasgutner, P. (2019). High prevalence and genetic diversity of Haemoproteus columbae (Haemosporida: Haemoproteidae) in feral pigeons Columba livia in Cape Town, South Africa. Parasitology Research.

Kleindorfer, S., Fessl, B., Peters, K., Anchundia, D. (2019) Field Guide. Resident land birds of Galapagos. Publication number 2223 of the Charles Darwin Foundation. ISBN: 978-9978-53-063-4.

Kleindorfer, S., Fessl, B., Peters, K., Anchundia, D. (2019) Guía de campo. Aves terrestres residentes de Galápagos. Publication number 2197 of the Charles Darwin Foundation. ISBN: 978-9978-53-062-7.

Loo, W.T., Dudaniec, R.Y., Kleindorfer, S., Cavanaugh, C.M. (2019) An inter-island comparison of Darwin’s finches reveals the impact of habitat, host phylogeny, and island on the gut microbiome. PLOS ONE

Loo, W.T., Dudaniec, R.Y., Loor, J.G., Kleindorfer, S., Cavanaugh, C.M. (2019) Host phylogeny, diet, and habitat differentiate the gut microbiomes of Darwin’s finches on Santa Cruz Island. Scientific Reports 9: 18781

Rivera, M., Cealie, M., Hauber, M.E., Kleindorfer, S., Liu, W. (2019) Neural activation in response to conspecific songs in zebra finch (Taeniopygia guttata) embryos and nestlings. NeuroReport 30(3): 217-221. DOI:

Sierro, J., Loretto, M.-C., Szipl, G., Massen, J. J. M., Bugnyar, T. (2019) Food calling in wild ravens (Corvus corax) revisited: Who is addressed? Ethology

Gallego-Abenza, M., Loretto, M.-C., Bugnyar, T. (2019) Decision time modulates social foraging success in wild common ravens, Corvus corax. Ethology, 00, 1-10

Richter, A., Sieber, A., Siebert, J., Miczajka-Rußmann, V. L., Zabel, J., Ziegler, D., Hecker, S., Frigerio, D. (2019) Storytelling for narrative approaches in citizen science: towards a generalized model. Journal of Science Communication, 18 (06), A02

Gallego-Abenza, M., Mathevon, N., Wheatcroft, D. (2019) Experience modulates an insect's response to anthropogenic noise. Behavioral Ecology, arz159. DOI:

Common, L. K., Dudaniec, R. Y., Colombelli-Négrel, D., Kleindorfer, S. (2019) Taxonomic shifts in Philornis larval behaviour and rapid changes in Philornis downsi Dodge & Aitken (Diptera: Muscidae): an invasive avian parasite on the Galápagos Islands. In: Life Cycle and Development of Diptera. DOI: 10.5772/intechopen.88854

Frigerio, D., Puehringer-Sturmayr, V., Neuböck-Hubinger, B., Gegendorfer, G., Kotrschal, K., Hirschenhauser, K. (2019) Monitoring public awareness about the endangered northern bald ibis: a case study involving primary school children as citizen scientists. PeerJ, 7, e7569

Szipl, G., Depenau, M., Kotrschal, K., Hemetsberger, J., Frigerio, D. (2019) Costs and benefits of social connectivity in juvenile greylag geese. Scientific Reports, 9, 12839

Hirschenhauser, K., Frigerio, D., Leighinger, V., Schenkenfelder, I., Neuböck-Hubinger, B. (2019) Primary pupils, science and a model bird species: evidence for the efficacy of extracurricular science education. PLoS ONE, 14 (7), e0220635

Kleindorfer, S., Custance, G., Peters, K. J., Sulloway, F. J. (2019) Introduced parasite changes host phenotype, mating signal and hybridization risk: Philornis downsi effects on Darwin's finch song. Proceedings of the Royal Society B: Biological Sciences, 286. DOI:

Frigerio, D., Hemetsberger, F., Wascher, C. A. F. (2019) How having a partner and a family changes the stress levels of greylag geese. Frontiers for Young Minds, 7 (68). DOI: 10.3389/frym.2019.00068

Langton, A., Kleindorfer, S. (2019) Minimum longevity and age-related male plumage in Darwin's finches on Floreana Island. Journal of Ornithology, 160 (2), 351-361

Peters, K. J., Evans, C., Aguirre, J. D., Kleindorfer, S. (2019) Genetic admixture predicts parasite intensity: evidence for increased hybrid performance in Darwin's tree finches. Royal Society Open Science, 6:181616. DOI:

Szipl, G., Loth, A., Wascher, C.A.F., Hemetsberger, J., Kotrschal, K., Frigerio, D. (2019) Parental behaviour and family proximity as key to gosling survival in Greylag Geese (Anser anser). Journal of Ornithology. DOI:


  Frigerio, D., Kotrschal, K., Fabro, C., Puehringer-Sturmayr, V., Iaiza, L., Hemetsberger, J., Mason, F., Sarnataro, C., Filacorda, S. (2018) Social context modulates digestive efficiency in greylag geese (Anser anser). Scientific reports, 8, 16498. DOI:

  Puehringer-Sturmayr, V., Gegendorfer, G., Rittenschober, J., Szipl, G., Frigerio, D. (2018). Involving pupils/citizens in long-term behavioural biology research: Lessons learnt and future perspectives. In: Heigl, F., Dörler, D., Ernst, M., editors. Proceedings of the Austrian Citizen Science Conference, 2018 Februray 1-3, Salzburg, Austria. Frontiers. p. 34-38. DOI:

  Szipl, G., Ringler, E., & Bugnyar, T. (2018). Attacked ravens flexibly adjust signalling behaviour according to audience composition. Proceedings of the Royal Society of London Series B: Biological Sciences, 285, (1880), 20180375.

  Ringler, E., Szipl, G., Harrigan, R. J., Bartl-Binder, P., Mangione, R., & Ringler, M. (2018). Hierarchical decision‐making balances current and future reproductive success. Molecular Ecology, 27(9), 2289–2301.

  Frigerio, D., Pipek, P., Kimmig, S., Winter, S., Melzheimer, J., Diblíková, L., et al. (2018). Citizen science and wildlife biology: Synergies and challenges. Ethology, 124, 365-377.

  Boeckle, M., Szipl, G., & Bugnyar, T. (2018). Raven food calls indicate sender's age and sex. Frontiers in Zoology, 15, 5.

  Wascher, C. A. F., Kotrschal, K., & Arnold, W. (2018). Free-living greylag geese adjust their heart rates and body core temperatures to season and reproductive context. Scientific Reports, 8(1), 2142.

  Puehringer-Sturmayr, V., Wascher, C. A. F., Loretto, M.-C., Palme, R., Stoewe, M., Kotrschal, K., & Frigerio, D. (2018). Seasonal differences of corticosterone metabolite concentrations and parasite burden in Northern Bald Ibis (Geronticus eremita): The role of affiliative interactions. PLoS ONE, e0191441.


Szipl, G., Ringler, E., Spreafico, M., & Bugnyar, T. (2017). Calls during agonistic interactions vary with arousal and raise audience attention in ravens. Frontiers in Zoology, 14, 57.

Loth, A., Frigerio, D., Kotrschal, K., & Szipl, G. (2017). Differential responses to gosling distress calls in parental and non-parental Greylag Geese. Journal of Ornithology, 159(2), 401-412.

Wascher, C. A. F., Hemetsberger, J., Kotrschal, K., & Frigerio, D. (2017). Leucocyte profiles and family size in fledgling Geylag Geese (Anser anser). Avian Biology Research10(4), 246–252.

Frigerio, D., Ludwig, S. C., Hemetsberger, J., Kotrschal, K., & Wascher, C. A. F. (2017). Social and environmental factors modulate leucocyte profiles in free-living Greylag geese (Anser anser). PeerJ, 5, e2792.

Loretto, M.-C., Schuster, R., Itty, C., Marchand, P., Genero, F., & Bugnyar, T. (2017). Fission-fusion dynamics over large distances in raven non-breeders. Scientific Reports, 7(1), 380.


Beck, K., Frigerio, D. & Loretto, M.-C. (2016). Der Kolkrabe – Schwarzfedriges Genie oder doch nur ein Unglücksbote? - Von Irrtümern und Mythen bis hin zur Forschung von Groß und Klein. Öko-L, 38(1), 3–11.

Frigerio, D., Cibulski, L., Ludwig, S. C., Campderrich, I., Kotrschal, K., & Wascher, C. A. F. (2016). Excretion patterns of coccidian oocysts and nematode eggs during the reproductive season in Northern Bald Ibis (Geronticus eremita). Journal of Ornithology, 157(3), 839–851.

Hirschenhauser, K., Frigerio, D. & Neuböck-Hubinger, B. (2016). Wirkungen außerschulischer Angebote im Sachunterricht: das Waldrapp-Projekt - Impact of external projects on teaching natural sciences in primary school: the Northern bald ibis case. In Transfer Forschung-Schule “Visible Didactics - Fachdidaktische Forschung trifft Praxis” (pp. 307–308). Bad Heilbrunn: Linkhard Verlag.

Loretto, M.-C., Schuster, R., & Bugnyar, T. (2016). GPS tracking of non-breeding ravens reveals the importance of anthropogenic food sources during their dispersal in the Eastern Alps. Current Zoology, 62(4), 337–344.


Loretto, M.-C., Reimann, S., Schuster, R., Graulich, D. M., & Bugnyar, T. (2015). Shared space, individually used: spatial behaviour of non-breeding ravens (Corvus corax) close to a permanent anthropogenic food source. Journal of Ornithology, 157(2), 439–450.

Szipl, G., Boeckle, M., Wascher, C. A. F., Spreafico, M., & Bugnyar, T. (2015). With whom to dine? Ravens’ responses to food-associated calls depend on individual characteristics of the caller. Animal Behaviour, 99(Supplement C), 33–42.


Cibulski, L., Wascher, C. A. F., Weiss, B. M., & Kotrschal, K. (2014). Familiarity with the experimenter influences the performance of Common ravens (Corvus corax) and Carrion crows (Corvus corone corone) in cognitive tasks. Behavioural Processes, 103, 129–137.

Dorn, S., Wascher, C. A. F., Möstl, E., & Kotrschal, K. (2014). Ambient temperature and air pressure modulate hormones and behaviour in Greylag geese (Anser anser) and Northern bald ibis (Geronticus eremita). Behavioural Processes, 108, 27–35.

Frigerio, D., Hemetsberger, J. & Kotrschal, K. (2014). Selbst nach 40 Jahre Forschung: Neues von den Grünauer Graugänsen - Von internationaler Grundlagenforschung bis zur Zusammenarbeit mit Volksschulkindern. Öko-L, 36(1), 3–13.

Hillemann, F., Bugnyar, T., Kotrschal, K., & Wascher, C. A. F. (2014). Waiting for better, not for more: corvids respond to quality in two delay maintenance tasks. Animal Behaviour, 90(Supplement C), 1–10.

Massen, J. J. M., Pašukonis, A., Schmidt, J., & Bugnyar, T. (2014). Ravens notice dominance reversals among conspecifics within and outside their social group. Nature Communications, 5, 3679.

Massen, J. J. M., Szipl, G., Spreafico, M., & Bugnyar, T. (2014). Ravens intervene in others’ bonding attempts. Current Biology, 24(22), 2733–2736.

Szipl, G., Boeckle, M., Werner, S. A. B., & Kotrschal, K. (2014). Mate Recognition and Expression of Affective State in Croop Calls of Northern Bald Ibis (Geronticus eremita). PLoS ONE, 9(2), e88265.

Szipl, G., & Bugnyar, T. (2014). Craving ravens: Individual “haa” call rates at feeding sites as cues to personality and levels of fission-fusion dynamics? Animal Behavior and Cognition, 1(3), 265–280.

Ujfalussy, D. J., Miklósi, Á., Bugnyar, T., & Kotrschal, K. (2014). Role of mental representations in quantity judgments by jackdaws (Corvus monedula). Journal of Comparative Psychology, 128(1), 11–20.


Bugnyar, T. (2013). Social cognition in ravens. Comparative Cognition & Behavior Reviews, 8, 1–12.

Frigerio, D., Gegendorfer, G. (2013). Exotisch oder heimisch? - Der Waldrapp aus dem Almtal. Groß und Klein erforschen die vom Aussterben bedrohten Vogelarten. Öko-L, 35(3), 3–13.

Guibert, F., Lumineau, S., Kotrschal, K., Möstl, E., Richard-Yris, M.-A., & Houdelier, C. (2013). Trans-generational effects of prenatal stress in quail. Proceedings. Biological Sciences, 280(1753).

Hirschenhauser, K., Gahr, M., & Goymann, W. (2013). Winning and losing in public: Audiences direct future success in Japanese quail. Hormones and Behavior, 63(4), 625–633.

Kotrschal, K. (2013). The Quest for Understanding Social Complexity. Quo Vadis, Behavioural Biology? Past, Present, and Future of an Evolving Science, 111(380), 77–88.

Mikolasch, S., Kotrschal, K., & Schloegl, C. (2013). Transitive inference in jackdaws. Behavioural Processes, 92, 113–117.

Schloegl, C., Waldmann, M. R., & Fischer, J. (2013). Understanding of and reasoning about object–object relationships in long-tailed macaques? Animal Cognition, 16(3), 493–507.

Teschke, I., Wascher, C. A. F., Scriba, M. F., von Bayern, A. M. P., Huml, V., Siemers, B., & Tebbich, S. (2013). Did tool-use evolve with enhanced physical cognitive abilities? Philosophical Transactions of the Royal Society of London. B, Biological Sciences, 368(1630), 20120418.

Wascher, C. A. F., & Bugnyar, T. (2013). Behavioral responses to inequity in reward distribution and working effort in crows and ravens. PLoS ONE, 8(2), e56885–e56885.

Weiss, B. M., & Scheiber, I. B. R. (2013). Long-term memory of hierarchical relationships in free-living greylag geese. Animal Cognition, 16(1), 91–97.


Bisenberger, A., Frigerio, D. (2012). Biene Maja zeigt uns ihre Welt - von einer projektidee zur angewandten Methodik tiergestützter Vermittlungsarbeit mit Kleintieren. Öko-L, 2, 29–35.

Boeckle, M., & Bugnyar, T. (2012). Long-term memory for affiliates in ravens. Current Biology, 22(9), 801–806.

Boeckle, M., Szipl, G., & Bugnyar, T. (2012). Who wants food? Individual characteristics in raven yells. Animal Behaviour, 84(5), 1123–1130.

Braun, A., & Bugnyar, T. (2012). Social bonds and rank acquisition in raven nonbreeder aggregations. Animal Behaviour, 84(6), 1507–1515.

Braun, A., Walsdorff, T., Fraser, O. N., & Bugnyar, T. (2012a). Socialized sub-groups in a temporary stable raven flock? Journal of Ornithology, 153(1), 97–104.

Braun, A., Walsdorff, T., Fraser, O. N., & Bugnyar, T. (2012b). Socialized sub-groups in a temporary stable Raven flock? Journal of Ornithology, 153(SUPPL. 1), 97–104.

Frigerio, D., Kotrschal, K., Millesi, E., & Hemetsberger, J. (2012). Children and Scientific Observations: Pupils Measuring Greylag Goose Behaviour. IJCDSE, 3, 871–876.

Dufour, V., Wascher, C. A. F., Braun, A., Miller, R., & Bugnyar, T. (2012). Corvids can decide if a future exchange is worth waiting for. Biology Letters, 8(2), 201–204.

Fraser, O. N., & Bugnyar, T. (2012). Reciprocity of agonistic support in ravens. Animal Behaviour, 83(1), 171–177.

Hirschenhauser, K., Spreitzer, K., Lepschy, M., Kotrschal, K., & Möstl, E. (2012). Excreted corticosterone metabolites differ between two galliform species, Japanese Quail and Chicken, between sexes and between urine and faecal parts of droppings. Journal of Ornithology, 153(4), 1179–1188.

Loretto, M.-C., Fraser, O. N., & Bugnyar, T. (2012). Ontogeny of Social Relations and Coalition Formation in Common Ravens (Corvus corax). International Journal of Comparative Psychology, 25, 180–194.

Schloegl, C., Schmidt, J., Boeckle, M., Weiss, B. M., & Kotrschal, K. (2012). Grey parrots use inferential reasoning based on acoustic cues alone. Proceedings. Biological Sciences, 279(1745), 4135–4142.

Schwab, C., Swoboda, R., Kotrschal, K., & Bugnyar, T. (2012). Recipients affect prosocial and altruistic choices in jackdaws, Corvus monedula. PLoS ONE, 7(4), e34922–e34922.

Wascher, C. A. F., Bauer, A. C., Holtmann, A. R., & Kotrschal, K. (2012). Environmental and social factors affecting the excretion of intestinal parasite eggs in graylag geese. Behavioral Ecology, 23(6), 1276–1283.

Wascher, C. A. F., Dufour, V., & Bugnyar, T. (2012). Carrion crows cannot overcome impulsive choice in a quantitative exchange task. Frontiers in Psychology, 3.

Wascher, C. A. F., Szipl, G., Boeckle, M., & Wilkinson, A. (2012). You sound familiar: carrion crows can differentiate between the calls of known and unknown heterospecifics. Animal Cognition, 15(5), 1015–1019.

Wascher, C. A. F., Weiss, B. M., Arnold, W., & Kotrschal, K. (2012). Physiological implications of pair-bond status in greylag geese. Biology Letters, 8(3), 347–350.


Beetz, A., Kotrschal, K., Turner, D. C., Hediger, K., Uvnäs-Moberg, K., & Julius, H. (2011). The effect of a real dog, toy dog and friendly person on insecurely attached children during a stressful task: An exploratory study. Anthrozoös, 24(4), 349–368.

Bugnyar, T. (2011). Knower-guesser differentiation in ravens: others’ viewpoints matter. Proceedings. Biological Sciences, 278(1705), 634–640.

Fraser, O. N., & Bugnyar, T. (2011). Ravens reconcile after aggressive conflicts with valuable partners. PLoS ONE, 6(3), e18118–e18118.

Kehmeier, S., Schloegl, C., Scheiber, I. B. R., & Weiss, B. M. (2011). Early development of gaze following into distant space in juvenile Greylag geese (Anser anser). Animal Cognition, 14(4), 477–485.

Kenward, B. E. N., Schloegl, C., Rutz, C., Weir, A. A. S., Bugnyar, T., & Kacelnik, A. (2011). On the evolutionary and ontogenetic origins of tool-oriented behaviour in New Caledonian crows (Corvus moneduloides). Biological Journal of the Linnean Society, 102(4), 870–877.

Mikolasch, S., Kotrschal, K., & Schloegl, C. (2011). African grey parrots (Psittacus erithacus) use inference by exclusion to find hidden food. Biology Letters, 7(6), 875–877.

Mikolasch, S., Kotrschal, K., & Schloegl, C. (2011). Is caching the key to exclusion in corvids? The case of carrion crows (Corvus corone corone). Animal Cognition, 15(1), 73–82.

Pika, S., & Bugnyar, T. (2011). The use of referential gestures in ravens (Corvus corax) in the wild. Nature Communications, 2, 560.

Scheiber, I. B. R., Hohnstein, A., Kotrschal, K., & Weiss, B. M. (2011). Juvenile greylag geese (Anser anser) discriminate between individual siblings. PLoS ONE, 6(8), e22853–e22853.

Schloegl, C. (2011). What you see is what you get—Reloaded: Can jackdaws (Corvus monedula) find hidden food through exclusion? Journal of Comparative Psychology, 125(2), 162–174.

Schmidt, J., Scheid, C., Kotrschal, K., Bugnyar, T., & Schloegl, C. (2011). Gaze direction – A cue for hidden food in rooks (Corvus frugilegus)? Behavioural Processes, 88(2), 88–93. doi.org

Wagner, J., Szipl, G., & Schwab, C. (2011). Successfully releasing Jackdaws, Corvus monedula: spatial dispersion and the fusion of social groups. Die Vogelwarte, 49, 163–174.

Wedl, M., Bauer, B., Gracey, D., Grabmayer, C., Spielauer, E., Day, J., & Kotrschal, K. (2011). Factors influencing the temporal patterns of dyadic behaviours and interactions between domestic cats and their owners. Behavioural Processes, 86(1), 58–67. doi.org

Weiß, B. M., Kotrschal, K., & Foerster, K. (2011). A longitudinal study of dominance and aggression in greylag geese (Anser anser). Behavioral Ecology, 22(3), 616–624.


Fraser, O. N., & Bugnyar, T. (2010a). Do ravens show consolation? Responses to distressed others. PLoS ONE, 5(5), e10605–e10605.

Fraser, O. N., & Bugnyar, T. (2010b). The quality of social relationships in ravens. Animal Behaviour, 79(4), 927–933.

Hemetsberger, J., Scheiber, I. B. R., Weiß, B. M., Frigerio, D., & Kotrschal, K. (2010). Influence of socially involved hand-raising on life history and stress responses in greylag geese. Interaction Studies, 11(3), 380–395.

Hirschenhauser, K., Weiß, B. M., Haberl, W., Möstl, E., & Kotrschal, K. (2010). Female androgen patterns and within-pair testosterone compatibility in domestic geese (Anser domesticus). General and Comparative Endocrinology, 165(2), 195–203. doi.org

Kotrschal, K., Scheiber, I. B. R., & Hirschenhauser, K. (2010). Individual performance in complex social systems: the greylag goose example BT - Animal Behaviour: Evolution and Mechanisms. In P. Kappeler (Ed.), Animal Behaviour: Evolution and Mechanisms (pp. 121–148). Berlin, Heidelberg: Springer Berlin Heidelberg.

Kralj-Fišer, S., Scheiber, I. B. R., Kotrschal, K., Weiß, B. M., & Wascher, C. A. F. (2010). Glucocorticoids enhance and suppress heart rate and behaviour in time dependent manner in greylag geese (Anser anser). Physiology & Behavior, 100(4), 394–400. doi.org

Kralj-Fišer, S., Weiß, B. M., & Kotrschal, K. (2010). Behavioural and physiological correlates of personality in greylag geese (Anser anser). Journal of Ethology, 28(2), 363–370.

Loretto, M.-C., Schloegl, C., & Bugnyar, T. (2010). Northern bald ibises follow others’ gaze into distant space but not behind barriers. Biology Letters, 6(1), 14 - 17.

Scheiber, I. B. R. (2010). Sozialverhalten im Lichte der Evolutionstheorie und Verhaltensbiologie. Evolution: Natur - Mensch - Gesellschaft.

Wascher, C. A. F., Fraser, O. N., & Kotrschal, K. (2010). Heart rate during conflicts predicts post-conflict stress-related behavior in Greylag geese. PLoS ONE, 5(12).

Wedl, M., Schöberl, I., Bauer, B., Day, J., & Kotrschal, K. (2010). Relational factors affecting dog social attraction to human partners. Interaction Studies, 11(3), 482–503.

Weiß, B. M., Kehmeier, S., & Schloegl, C. (2010). Transitive inference in free-living greylag geese, Anser anser. Animal Behaviour, 79(6), 1277–1283. doi.org

Weiss, B. M., Kotrschal, K., Möstl, E., & Hirschenhauser, K. (2010). Social and life-history correlates of hormonal partner compatibility in Greylag geese (Anser anser). Behavioral Ecology, 21(1), 138–143.

Weiß, B. M., Kotrschal, K., Möstl, E., & Hirschenhauser, K. (2010). Social and life-history correlates of hormonal partner compatibility in greylag geese (Anser anser). Behavioral Ecology, 21(1), 138–143.


Bertin, A., Hirschenhauser, K., & Kotrschal, K. (2009). Trans-Generational Influence of Human Disturbances in Japanese Quail: Egg Quality Influences Male Social and Sexual Behaviour. Ethology, 115(9), 879–887.

Bertin, A., Richard-Yris, M. A., Houdelier, C., Richard, S., Lumineau, S., Kotrschal, K., & Möstl, E. (2009). Divergent selection for inherent fearfulness leads to divergent yolk steroid levels in quail. Behaviour, 146(6), 757–770. doi.org

Day, J.E.L., Kergoat, S., Kotrschal, K. (2009). Do pets influence the quantity and choice of food offered to them by their owners: lessons from other animals and the pre-verbal human infant? CAB Reviews: Perspectives in Agriculture, Veterinary Science, Nutrition and Natural Resources, 4(42), 1–12.

Kotrschal, K. (2009). Die evolutionäre Theorie der Mensch-Tier-Beziehung. OTTERSTEDT, C./ROSENBERGER, M.(HG), Gefährten--Konkurrenten--Verwandte. Die Mensch-Tier-Beziehung Im Wissenschaftlichen Diskurs, Göttingen, 55–77.

Kotrschal, K., Schöberl, I., Bauer, B., Thibeaut, A.-M., & Wedl, M. (2009). Dyadic relationships and operational performance of male and female owners and their male dogs. Behavioural Processes, 81(3), 383–391. doi.org

Scheiber, I. B. R., Kotrschal, K., & Weiss, B. M. (2009a). Benefits of family reunions: Social support in secondary Greylag goose families. Hormones and Behavior, 55(1), 133–138.

Scheiber, I. B. R., Kotrschal, K., & Weiss, B. M. (2009b). Serial agonistic attacks by Greylag goose families, Anser anser, against the same opponent. Animal Behaviour, 77(5), 1211–1216.

Schloegl, C., Bugnyar, T., & Aust, U. (2009). Exclusion performances in non-human animals: from pigeons to chimpanzees and back again. Rational Animals, Irrational Humans. Keio University Press, Tokyo, 217–234.

Schloegl, C., Dierks, A., Gajdon, G. K., Huber, L., Kotrschal, K., & Bugnyar, T. (2009). What you see is what you get? Exclusion performances in ravens and keas. PLoS ONE, 4(8).

Wascher, C. A. F., Scheiber, I. B. R., Weiß, B. M., & Kotrschal, K. (2009). Heart rate responses to agonistic encounters in greylag geese, Anser anser. Animal Behaviour, 77(4), 955–961. doi.org

Wedl, M., Bauer, B., Dittami, J., Schöberl, I., & Kotrschal, K. (2009). Effects of personality and sex on behavioral patterns in human--dog dyads. Journal of Veterinary Behavior: Clinical Applications and Research, 4(2), 88–89.

Wedl, M., & Kotrschal, K. (2009). Social and Individual Components of Animal Contact in Preschool Children. Anthrozoös, 22(4), 383–396.


Bertin, A., Richard-Yris, M.-A., Houdelier, C., Lumineau, S., Möstl, E., Kuchar, A., … Kotrschal, K. (2008). Habituation to humans affects yolk steroid levels and offspring phenotype in quail. Hormones and Behavior, 54(3), 396–402. doi.org

Bókony, V., Garamszegi, L. Z., Hirschenhauser, K., & Liker, A. (2008). Testosterone and melanin-based black plumage coloration: a comparative study. Behavioral Ecology and Sociobiology, 62(8).

Bugnyar, T. (2008). Animal Cognition: Rooks Team up to Solve a Problem. Current Biology, 18(12), R530–R532. doi.org

Garamszegi, L. Z., Hirschenhauser, K., Bókony, V., Eens, M., Hurtrez‐Boussès, S., Møller, A. P., … Wingfield, J. C. (2008). Latitudinal Distribution, Migration, and Testosterone Levels in Birds. The American Naturalist, 172(4), 533–546.

Hirschenhauser, K., Canário, A., Ros, A., Taborsky, M., & Oliveira, R. (2008). Social context may affect urinary excretion of 11-ketotestosterone in African cichlids. Behaviour, 145(10), 1367–1388. doi.org

Hirschenhauser, K., Wittek, M., Johnston, P., & Möstl, E. (2008). Social context rather than behavioral output or winning modulates post-conflict testosterone responses in Japanese quail (Coturnix japonica). Physiology & Behavior, 95(3), 457–463. doi.org

Kotrschal, K. (2008). Angepasst woran? Vom Mängel- zum Konfliktwesen. In C. Schmidinger, H., Sedmak (Ed.), Der Mensch - Ein Mängelwesen? Endlichkeit - Kompensation - Entwicklung (pp. 55–65). Darmstadt: WBG.

Kotrschal, K. (2008). Geist als Anspannung. In S. Appel, K., Weber, H.P., Langthaler, R., Müller (Ed.), Naturalisierung des Geistes? Beiträge zur gegenwärtigen Debatte um den Geist (pp. 84–92). Würzburg: Königshaus und Neumann.

Range, F., Bugnyar, T., & Kotrschal, K. (2008). The performance of ravens on simple discrimination tasks: a preliminary study. Acta Ethologica, 11(1), 34–41.

Scheiber, I. B. R., Weiss, B. M., Hirschenhauser, K., Wascher, C. A. F., Nedelcu, I. T., & Kotrschal, K. (2008). Does “relationship intelligence” make big brains in birds? The Open Biology Journal, 1, 6–8.

Scheid, C., & Bugnyar, T. (2008). Short-term observational spatial memory in Jackdaws (Corvus monedula) and Ravens (Corvus corax). Animal Cognition, 11(4), 691–698.

Scheid, C., Schmidt, J., & Noë, R. (2008). Distinct patterns of food offering and co-feeding in rooks. Animal Behaviour, 76(5), 1701–1707. doi.org

Schloegl, C., Kotrschal, K., & Bugnyar, T. (2008a). Do common ravens (Corvus corax) rely on human or conspecific gaze cues to detect hidden food? Animal Cognition, 11(2), 231–241.

Schloegl, C., Kotrschal, K., & Bugnyar, T. (2008b). Modifying the object-choice task: Is the way you look important for ravens? Behavioural Processes, 77(1), 61–65. doi.org

Schloegl, C., Schmidt, J., Scheid, C., Kotrschal, K., & Bugnyar, T. (2008). Gaze following in non-human animals: the corvid example. Animal Behaviour: New Research, 73–92.

Scholl, S., Grall, G., Petzl, V., Röthler, M., Slotta-Bachmayr, L., & Kotrschal, K. (2008). Behavioural effects of goats on disabled persons. Therapeutic Communities, 29(3), 297–309.

Schwab, C., Bugnyar, T., & Kotrschal, K. (2008). Preferential learning from non-affiliated individuals in jackdaws (Corvus monedula). Behavioural Processes, 79(3), 148–155.

Schwab, C., Bugnyar, T., Schloegl, C., & Kotrschal, K. (2008). Enhanced social learning between siblings in common ravens, Corvus corax. Animal Behaviour, 75(2), 501–508.

Scott, A., Hirschenhauser, K., Bender, N., Oliveira, R., Earley, R., Sebire, M., … Canario, A. (2008). Non-invasive measurement of steroids in fish-holding water: important considerations when applying the procedure to behaviour studies. Behaviour, 145(10), 1307–1328. doi.org

Stöwe, M., Bugnyar, T., Schloegl, C., Heinrich, B., Kotrschal, K., & Möstl, E. (2008). Corticosterone excretion patterns and affiliative behavior over development in ravens (Corvus corax). Hormones and Behavior, 53(1), 208–216. doi.org

Wascher, C. A. F., Arnold, W., & Kotrschal, K. (2008). Heart rate modulation by social contexts in Greylag geese (Anser anser). Journal of Comparative Psychology, 122(1), 100–107.

Wascher, C. A. F., Scheiber, I. B. R., & Kotrschal, K. (2008). Heart rate modulation in bystanding geese watching social and non-social events. Proceedings. Biological Sciences, 275(1643), 1653–1659.

Weiß, B. M., Kotrschal, K., & Frigerio, D. (2008). Birds of a feather stay together: extended family bonds, clan structures and social support in greylag geese. In Family relations issues and challenges (pp. 69–88). Nova Science Publishers.

Weiß, B. M., Poggemann, K., Olek, K., Foerster, K., & Hirschenhauser, K. (2008). Isolation and characterization of microsatellite marker loci in the greylag goose (Anser anser). Molecular Ecology Resources, 8(6), 1411–1413.


Bugnyar, T. (2007). An integrative approach to the study of "theory-of-mind"-like abilities in ravens. 動物心理学研究, 57(1), 15–27.

Bugnyar, T., Schwab, C., Schloegl, C., Kotrschal, K., & Heinrich, B. (2007). Ravens Judge Competitors through Experience with Play Caching. Current Biology, 17(20), 1804–1808. doi.org

Bugnyar, T., Stöwe, M., & Heinrich, B. (2007). The ontogeny of caching in ravens, Corvus corax. Animal Behaviour, 74(4), 757–767. doi.org

Christian, Kotrschal, K., & Thomas. (2007). Lektionen von Rabenvögeln und Gänsen. Instinkt oder Verstand? Biologie in Unserer Zeit, 37(6), 366–374.

Kralj-Fišer, S., Scheiber, I. B. R., Blejec, A., Moestl, E., & Kotrschal, K. (2007). Individualities in a flock of free-roaming greylag geese: Behavioral and physiological consistency over time and across situations. Hormones and Behavior, 51(2), 239–248. doi.org

Scheid, C., Range, F., & Bugnyar, T. (2007). When, What, and Whom to Watch? Quantifying Attention in Ravens (Corvus corax) and Jackdaws (Corvus monedula). Journal of Comparative Psychology, 121(4), 380–386.

Schloegl, C., Kotrschal, K., & Bugnyar, T. (2007). Gaze following in common ravens, Corvus corax: ontogeny and habituation. Animal Behaviour, 74(4), 769–778. doi.org

Stöwe, M., & Kotrschal, K. (2007). Behavioural phenotypes may determine whether social context facilitates or delays novel object exploration in ravens (Corvus corax). Journal of Ornithology, 148(2), 179–184.

Zoufal, K., Fritz, J., Bichler, M., Kirbauer, M., Markut, T., Meran, I., … Kotrschal, K. (2007). Feeding ecology of the Northern Bald Ibis in different habitat types: an experimental field study with hand-raised individuals.


Bugnyar, T., & Heinrich, B. (2006). Pilfering ravens, Corvus corax, adjust their behaviour to social context and identity of competitors. Animal Cognition, 9(4), 369–376.

Hirschenhauser, K., & Oliveira, R. F. (2006). Social modulation of androgens in male vertebrates: meta-analyses of the challenge hypothesis. Animal Behaviour, 71(2), 265–277.

Kotrschal, K., Hemetsberger, J., & Weiß, B. M. (2006). Homosociality in male Greylag geese (Anser anser): making the best of a bad situation. Homosexual Behaviour in Animals: An Evolutionary Perspective. Cambridge University Press Cambridge.

Range, F., Bugnyar, T., Schlögl, C., & Kotrschal, K. (2006). Individual and sex differences in learning abilities of ravens. Behavioural Processes, 73(1), 100–106.

Sorato, E., & Kotrschal, K. (2006). Hormonal and behavioural symmetries between the sexes in the Northern bald ibis. General and Comparative Endocrinology, 146(3), 265–274.

Stöwe, M., Bugnyar, T., Heinrich, B., & Kotrschal, K. (2006). Effects of Group Size on Approach to Novel Objects in Ravens (Corvus corax). Ethology, 112(11), 1079–1088.

Stöwe, M., Bugnyar, T., Loretto, M.-C., Schloegl, C., Range, F., & Kotrschal, K. (2006). Novel object exploration in ravens (Corvus corax): Effects of social relationships. Behavioural Processes, 73(1), 68–75. doi.org


Bugnyar, T., Stöwe, M., & Heinrich, B. (2005). Ravens, Corvus corax, follow gaze direction of humans around obstacles. Proceedings. Biological Sciences, 271(1546).

Heinrich, B., & Bugnyar, T. (2005). Testing Problem Solving in Ravens: String-Pulling to Reach Food. Ethology, 111(10), 962–976.

Hemetsberger, J. (2005). Frühe Brutnachweise bei Steinlerche Ammomanes isabellina, Saharasteinschmätzer Oenanthe leucopyga, Akaziendrossling Turdoides fulvus und Hausammer Emberiza striolata in der zentralen Sahara Algeriens. Vogelwarte, 271.

Hirschenhauser, K., & Frigerio, D. (2005). 5. Hidden Patterns of Male Sex Hormones and Behaviour Vary with Life History. In G. R. L. Anolli , S. Duncan , M. S. Magnusson (Ed.), The hidden structure of social interaction. Amsterdam: Citeseer.

Hirschenhauser, K., Kotrschal, K., & Möstl, E. (2005). Synthesis of Measuring Steroid Metabolites in Goose Feces. Annals of the New York Academy of Sciences, 1046(1), 138–153.

Niall Daisley, J., Bromundt, V., Möstl, E., & Kotrschal, K. (2005). Enhanced yolk testosterone influences behavioral phenotype independent of sex in Japanese quail chicks Coturnix japonica. Hormones and Behavior, 47(2), 185–194. doi.org

Scheiber, I. B. R., Kralj, Si., & Kotrschal, K. (2005). Sampling Effort/Frequency Necessary to Infer Individual Acute Stress Responses from Fecal Analysis in Greylag Geese (Anser anser). Annals of the New York Academy of Sciences, 1046(1), 154–167.

Scheiber, I. B. R., Weiss, B. M., Frigerio, D., & Kotrschal, K. (2005). Active and passive social support in families of Greylag geese (Anser anser). Behaviour, 142, 1535–1557.

Wei, B. M., Möstl, E., & Hirschenhauser, K. (2005). Within-pair testosterone compatibility as a currency for pairbond quality in greylag geese? Hormones and Behavior, 48(1), 133–134.


Bugnyar, T., & Kotrschal, K. (2004). Leading a conspecific away from food in ravens (Corvus corax)? Animal Cognition, 7(2), 69–76.

Bugnyar, T., Stöwe, M., & Heinrich, B. (2004). Ravens, Corvus corax, follow gaze direction of humans around obstacles. Proceedings of the Royal Society of London. Series B: Biological Sciences, 271(1546).

Frigerio, D., Dittami, J., Möstl, E., & Kotrschal, K. (2004). Excreted corticosterone metabolites co-vary with ambient temperature and air pressure in male Greylag geese (Anser anser). General and Comparative Endocrinology, 137(1), 29–36. doi.org

Frigerio, D., Hirschenhauser, K., Möstl, E., Dittami, J., & Kotrschal, K. (2004). Experimentally elevated testosterone increases status signalling in male Greylag geese (Anser anser). Acta Ethologica, 7(1), 9–18.

Goymann, W., Moore, I. T., Scheuerlein, A., Hirschenhauser, K., Grafen, A., & Wingfield, J. C. (2004). Testosterone in Tropical Birds: Effects of Environmental and Social Factors. The American Naturalist, 164(3), 327–334.

Hemetsberger, J., & Petutschnig, B. (2004). Zur Avifauna Jordaniens.

Hirschenhauser, K., Taborsky, M., Oliveira, T., Canàrio, A. V. M., & Oliveira, R. F. (2004). A test of the “challenge hypothesis” in cichlid fish: simulated partner and territory intruder experiments. Animal Behaviour, 68(4), 741–750. doi.org

Kotrschal, K., Bromundt, V., & Föger, B. (2004). Faktor Hund: eine sozio-ökonomische Bestandsaufnahme der Hundehaltung in Österreich. Czernin.

Scheiber, I. B. (2004). Social Support As The Key Factor For Individual Success In Greylag Geese (Anser anser)? Hormones and Behavior, 46(1), 121.

Weiß, B. M., & Kotrschal, K. (2004). Effects of Passive Social Support in Juvenile Greylag Geese (Anser anser): A Study from Fledging to Adulthood. Ethology, 110(6), 429–444.


Frigerio, D., Weiss, B., Dittami, J., & Kotrschal, K. (2003). Social allies modulate corticosterone excretion and increase success in agonistic interactions in juvenile hand-raised graylag geese (Anser anser). Canadian Journal of Zoology, 81(10), 1746–1754.

Hackl, R., Bromundt, V., Daisley, J., Kotrschal, K., & Möstl, E. (2003). Distribution and origin of steroid hormones in the yolk of Japanese quail eggs (Coturnix coturnix japonica). Journal of Comparative Physiology B, 173(4), 327–331.

Hirschenhauser, K., Winkler, H., & Oliveira, R. F. (2003). Comparative analysis of male androgen responsiveness to social environment in birds: the effects of mating system and paternal incubation. Hormones and Behavior, 43(4), 508–519. doi.org

Kotrschal, K. (2003). Das Ende der Beliebigkeit — zum evolutionär-biologischen Fundament des Menschen. Österreichische Zeitschrift Für Soziologie, 28(2), 21–43.

Kotrschal, K. (2003). Im Egoismus vereint? (2nd ed.). Fürth: Filander Verlag.

Kotrschal, K., & Ortbauer, B. (2003). Behavioral effects of the presence of a dog in a classroom. Anthrozoös, 16(2), 147–159.


Bugnyar, T., & Kotrschal, K. (2002a). Observational learning and the raiding of food caches in ravens, Corvus corax: is it “tactical” deception? Animal Behaviour, 64(2), 185–195.

Bugnyar, T., & Kotrschal, K. (2002b). Scrounging Tactics in Free-Ranging Ravens, Corvus corax. Ethology, 108(11), 993–1009.

Fritz, J., & Kotrschal, K. (2002). On avian imitation: cognitive and ethological perspectives. Imitation in Animals and Artifact, 133–155.

Hirschenhauser, K., Frigerio, D., Grammer, K., & Magnusson, M. S. (2002). Monthly Patterns of Testosterone and Behavior in Prospective Fathers. Hormones and Behavior, 42(2), 172–181. doi.org

Kijne, M., & Kotrschal, K. (2002). Neophobia affects choice of food-item size in group-foraging common ravens (Corvus corax). Acta Ethologica, 5(1), 13–18.

Kotrschal, K. (2002). Eine integrative, systematische Sicht sozialer. In U. Ganßloser (Ed.), Soziale Mechanismen. Fürth: Filander Verlag.

Pfeffer, K., Fritz, J., & Kotrschal, K. (2002). Hormonal correlates of being an innovative greylag goose, Anser anser. Animal Behaviour, 63(4), 687–695. doi.org

Tintner, A., & Kotrschal, K. (2002). Early social influence on nestling development in Waldrapp ibis (Geronticus eremita). Zoo Biology, 21(5), 467–480.


Bugnyar, T., Kijne, M., & Kotrschal, K. (2001). Food calling in ravens: are yells referential signals? Animal Behaviour, 61(5), 949–958.

Bugnyar, T., & Kotrschal, K. (2001). Movement coordination and signalling in ravens (Corvus corax): an experimental field study. Acta Ethologica, 3(2), 101–109.

Frigerio, D., Moestl, E., & Kotrschal, K. (2001). Excreted Metabolites of Gonadal Steroid Hormones and Corticosterone in Greylag Geese (Anser anser) from Hatching to Fledging. General and Comparative Endocrinology, 124(2), 246–255. doi.org

Frigerio, D., Weiss, B. M., & Kotrschal, K. (2001). Spatial proximity among adult siblings in Greylag geese (Anser anser): evidence for female bonding? Acta Ethologica, 3(2), 121–125.

Fritz, J. (2001). Der intelligente Kumpan am Beispiel der Graugänse. In H. . Kotrschal, Kurt; Müller, G.; Winkler (Ed.), Konzepte der Verhaltensforschung. Konrad Lorenz und die Folgen. Fürth: Filander Verlag.

Hemetsberger, J. (2001). Die Entwicklung der Grünauer Graugansschar seit 1973. (Kotrschal,K., Müller, G.,Winkler, Ed.), Konrad Lorenz und seine verhaltensbiologischen Konzepte aus heutiger Sicht. Fürth: Filander Verlag.

Kotrschal, K. (2001). Das Rousseausche Vorurteil vom „edlen Wilden “: Die Wurzeln der Ethik von Konrad Lorenz. (G. M. & H. W. K. Kotrschal, Ed.). Fürth: Filander Verlag.

Kotrschal, K. (2001). Der Beitrag von Österreich zur Wissenschaft der Biologie vom Verhalten und zum evolutionären Bild des Menschen. In K. Acham (Ed.), Cognitio Humana. Zur Geistesgeschichte Österreichs.

Kotrschal, K., Bugnyar, T., & Stöwe, M. (2001). Kognition und Neophobie bei Raben. Charandrius, 3, 127–135.

Mostl, E., Spendier, H., & Kotrschal, K. (2001). Concentration of immunoreactive progesterone and androgens in the yolk of hens’ eggs (Gallus domesticus). Wiener Tierarztliche Monatsschrift, 88(3), 62–65.

Older Publications

Fritz, J., Bisenberger, A., & Kotrschal, K. (2000). Stimulus enhancement in greylag geese: socially mediated learning of an operant task. Animal Behaviour, 59(6), 1119–1125. doi.org

Hemetsberger, J. (2000). Erste Ergebnisse des Jahres 1999. Vogelkundliche Nachrichten Oberösterreich - Naturschutz Aktuell, 8(1), 19–27.

Hirschenhauser, K., Möstl, E., Péczely, P., Wallner, B., Dittami, J., & Kotrschal, K. (2000). Seasonal Relationships between Plasma and Fecal Testosterone in Response to GnRH in Domestic Ganders. General and Comparative Endocrinology, 118(2), 262–272. doi.org

Hirschenhauser, K., Möstl, E., Wallner, B., Dittami, J., & Kotrschal, K. (2000). Endocrine and Behavioural Responses of Male Greylag Geese (Anser anser) to Pairbond Challenges during the Reproductive Season. Ethology, 106(1), 63–77.

Kotrschal, K. (2000). Taste(s) and olfaction(s) in fish: a review of spezialized sub-systems and central integration. Pflügers Archiv - European Journal of Physiology, 439(7), R178–R180.

Kotrschal, K., Dittami, J., Hirschenhauser, K., Möstl, E., & Peczely, P. (2000). Effects of physiological and social challenges in different seasons on fecal testosterone and corticosterone in male domestic geese (Anser domesticus). Acta Ethologica, 2(2), 115–122.

Fritz, J., & Kotrschal, K. (1999). Social learning in common ravens,Corvus corax. Animal Behaviour, 57(4), 785–793. doi.org

Hirschenhauser, K., Möstl, E.,Kotrschal, K. (1999). Faecal analysis of androgens and its relevance for the study of social behaviour and reproductive success in Greylag geese Anser anser. Ethology, 34, 206.

Hirschenhauser, K., Mostl, E., & Kotrschal, K. (1999). Within-pair testosterone covariation and reproductive output in Greylag Geese Anser anser. Ibis, 141(4), 577–586.

Hirschenhauser, K., Möstl, E., & Kotrschal, K. (1999). Seasonal Patterns of Sex Steroids Determined from Feces in Different Social Categories of Greylag Geese (Anser anser). General and Comparative Endocrinology, 114(1), 67–79. doi.org

Kotrschal, Kurt, Hirschenhauser, Katharina, Möstl, E., Peczely, P. (1999). Testosterone and corticosterone determination from feces in geese yields biologically relevant results. Ethology, 34, 207.

Kotrschal, K. (1998). Sensory Systems. In M. Horn, M., Martin, K., Chotkowski (Ed.), Intertidal Fishes (pp. 126–142). San Diego: Academic Press.

Kotrschal, K., Hirschenhauser, K., & Möstl, E. (1998). The relationship between social stress and dominance is seasonal in greylag geese. Animal Behaviour, 55(1), 171–176. doi.org

Kotrschal, K., Van Staaden, M. J., & Huber, R. (1998). Fish Brains: Evolution and Anvironmental Relationships. Reviews in Fish Biology and Fisheries, 8(4), 373–408.

Frigerio, D., Hirschenhauser, K., Wallner, B., & Kotrschal, K. (1997). Experimentally increased testosterone affects behaviour and corticosterone in male greylag geese. Advances in Ethology, 32, 59.

Hirschenhauser, K., Dittami, J., Möstl, E., Wallner, B., & Kotrschal, K. (1997). Testosterone and behaviour during different phases of reproduction in greylag geese (Anser anser). Advances in Ethology, 32, 61.

Kotrschal, K. (1997). Die Konrad Lorenz Forschungsstelle in Grünau. Vogelkundliche Nachrichten Oberösterreich - Naturschutz Aktuell, 4, 31–33.

Kotrschal, K., Krautgartner, W.-D., & Hansen, A. (1997). Ontogeny of the Solitary Chemosensory Cells in the Zebrafish, Danio rerio. Chemical Senses, 22(2), 111–118.

Weiß, B. M., & Kotrschal, K. (1997). Humans as social allies: interspecific social support in greylag geese (Anser anser). Advances in Ethology; Blackwell Sciences, 32, 150.

Kotrschal, K. (1996). Solitary chemosensory cells: why do primary aquatic vertebrates need another taste system? Trends in Ecology & Evolution, 11(3), 110–114. doi.org

Kotrschal, K. (1996). Solitary chemosensory cells: why do primary aquatic vertebrates need another taste system? Trends in Ecology & Evolution, 11(3), 110–114. doi.org

Kotrschal, K., Peters, R. C., & Døving, K. B. (1996). Chemosensory and tactile nerve responses from the anterior dorsal fin of a rockling, Gaidropsarus vulgaris (Gadidae, Teleostei). Primary Sensory Neuron, 1, 297–310.

Drack, G., & Kotrschal, K. (1995). Aktivitätsmuster und Spiel von freilebenden Kolkraben Corvus corax im inneren Almtal/Oberösterreich. Monticula, 7, 159–174.

Kotrschal, K. (1995). Ecomorphology of solitary chemosensory cell systems in fish: a review. Environmental Biology of Fishes, 44(1), 143–155.

Kotrschal, K., & Essler, H. (1995). Goals and approaches in the analysis of locomotion in fish, with a focus on laboratory studies. Reviews in Fisheries Science, 3(2), 171–200.

Essler, H., & Kotrschal, K. (1994). High resolution analysis of swim path patterns of intact and olfaction-deprived minnows (Phoxinus phoxinus) stimulated with food and potential predator odour. Journal of Fish Biology, 45(4), 555–567.

Kotrschal, K., Peters, R., & Atema, J. (1994). Zoology : analysis of complex systems : ZACS. Zoologische Jahrbücher. Abteilung für allgemeine Zoologie und Physiologie der Tiere (Vol. 97). G. Fischer.

Kotrschal, K., Hemetsberger, J., & Dittami, J. (1993). Food exploitation by a winter flock of greylag geese: behavioral dynamics, competition and social status. Behavioral Ecology and Sociobiology, 33(5), 289–295.

Kotrschal, K., Peters, R., & Atema, J. (1993). Sampling and behavioral evidence for mucus detection in a unique chemosensory organ: the anterior dorsal fin in rocklings (Ciliata mustela: Gadidae: Teleostei). Zoologische Jahrbücher. Abteilung Für Allgemeine Zoologie Und Physiologie Der Tiere, 97(1), 47–67.

Kotrschal, K., Whitear, M., & Finger, T. E. (1993). Spinal and facial innervation of the skin in the gadid fish Ciliata mustela (Teleostei). The Journal of Comparative Neurology, 331(3), 407–417.

Kotrschal, K. (1992). Quantitative scanning electron microscopy of solitary chemoreceptor cells in cyprinids and other teleosts. Environmental Biology of Fishes, 35(3), 273–282.

Kotrschal, K., Hemetsberger, J., & Dittami, J. (1992). Vigilance in a flock of semi-tame Greylag Geese Answer answer in response to approaching eagles Haliaeetus albicilla and Aquila chrysaetos. Wildfowl, 43(43), 215–219.

Kotrschal, K., & Palzenberger, M. (1992). Neuroecology of cyprinids: comparative, quantitative histology reveals diverse brain patterns BT - Environmental biology of European cyprinids: Papers from the workshop on “The Environmental Biology of Cyprinids” held at the University of Salzburg, Austr. In W. Wieser, F. Schiemer, A. Goldschmidt, & K. Kotrschal (Eds.) (pp. 135–152). Dordrecht: Springer Netherlands.

Mausz, B., Dittami, J., Kotrschal, K. (1992). Triumphgeschrei und Aggression bei der Graugans (Anser anser). Ökologie Der Vögel, 14, 165–172.

Finger, T. E., Drake, S. K., Kotrschal, K., Womble, M., & Dockstader, K. C. (1991). Postlarval growth of the peripheral gustatory system in the channel catfish, Ictalurus punctatus. The Journal of Comparative Neurology, 314(1), 55–66.

Kotrschal, K. (1991). Solitary chemosensory cells — taste, common chemical sense or what? Reviews in Fish Biology and Fisheries, 1(1), 3–22.

Peters, R. C., Kotrschal, K., & Krautgartner, W.-D. (1991). Solitary chemoreceptor cells of Ciliata mustela (Gadidae, Teleostei) are tuned to mucoid stimuli. Chemical Senses, 16(1), 31–42.

Kotrschal, K. (1989). Trophic ecomorphology in eastern Pacific blennioid fishes: character transformation of oral jaws and associated change of their biological roles. Environmental Biology of Fishes, 24(3), 199–218.

Kotrschal, B. K. (1988). Evolutionary patterns in tropical marine reef fish feeding. Journal of Zoological Systematics and Evolutionary Research, 26(1), 51–64.

Kotrschal, K., & Whitear, M. (1988). Chemosensory anterior dorsal fin in rocklings (Gaidropsarus and Ciliata, Teleostei, Gadidae): Somatotopic representation of the ramus recurrens facialis a revealed by transganglionic transport of HRP. The Journal of Comparative Neurology, 268(1), 109–120.

Lindquist, D. G., & Kotrschal, K. M. (1987). The Diets in Four Pacific Tube Blennies (Acanthemblemaria: Chaenopsidae): Lack of Ecological Divergence in Syntopic Species. Marine Ecology, 8(4), 327–335.

Kotrschal, K., & Lindquist, D. G. (1986). The Feeding Apparatus in four Pacific Tube Blennies (Te/eos fei: Chaenopsidae): Marine Ecology, 7(3), 241–253.

Goldschmid, A., & Kotrschal, K. (1985). Morphological and functional adaptations in different feeding types of blennies (Blenniidae, Perciformes, Teleostei). Forts Zool, 30, 241–244.