Frigerio, D., Colombelli-Négrel, D., Common, L.K., Katsis, A.C. and Kleindorfer, S. (2024) When parasites damage birds’ beaks, the whole environment can change. Front. Young Minds 12:1272047. doi: 10.3389/frym.2024.1272047

Naikatini, A.N., Keppel, G., Brodie, G. and Kleindorfer, S., 2024. Elevational differences in territory defence response in native (endemic and non-endemic) forest birds on Viti Levu Island, Fiji. New Zealand Journal of Zoology, pp.1-16.

Akçay C., Colombelli-Négrel D., Kleindorfer S. (2024) Buzzes are used as signals of aggressive intent in Darwin’s finches, Biological Journal of the Linnean Society, 2024;, blad152,

Frigerio D, Kleindorfer S, Krejci J, Schuster R, Kotrschal K, Loretto MC and Puehringer-Sturmayr V (2024) Saving Birds From Extinction by Discovering Their Habitats. Front. Young Minds. 12:1210462. doi: 10.3389/frym.2023.1210462

Kleindorfer S, Brouwer L, Hauber ME, Evans CE, Teunissen N, Peters A, Louter M, Webster M, Katsis AC, Sulloway FJ, Common LK, Austin VA, Colombelli-Négrel D (in press). Nestling begging calls resemble maternal vocal signatures when mothers call slowly to embryos. American Naturalist.


Colombelli-Négrel D, Akcay C, Kleindorfer S (2023). Darwin’s finches in human-altered environments sing common song types and are more aggressive. Frontiers in Ecology and Evolution 11.

Colombelli-Négrel, D., Sach, I.Z., Hough, I., Hodgson, J.C., Daniels, C.B. and Kleindorfer, S., 2023. Koalas showed limited behavioural response and no physiological response to drones. Applied Animal Behaviour Science, p.105963.

Common, L. K., Kleindorfer, S., Colombelli-Négrel, D., & Dudaniec, R. Y. (2023). Genetics reveals shifts in reproductive behaviour of the invasive bird parasite Philornis downsi collected from Darwin’s finch nests. Biological Invasions, 25, 563-581.

Frigerio D, Hemetsberger F, Sumasgutner P, Kotrschal K, Kleindorfer S and Hemetsberger J (2023) How Warming Temperatures Affect Breeding Behavior in Graylag Geese. Front. Young Minds. 11:922688. doi: 10.3389/frym.2023.922688

Frigerio D., Hohl A., Puehringer-Sturmayr V., Colombelli-Négrel D., Kleindorfer S. (2023) A direct personal experience of science and nature changes intended behaviours for conservation, Transactions of the Royal Society of South Australia, DOI:  10.1080/03721426.2023.2264366

Headland T, Colombelli-Négrel D, Callaghan CT, Sumasgutner SC, Kleindorfer S, Sumasgutner P: Smaller Australian raptors have greater urban tolerance. Scientific Reports 2023, 13(1):11559.

Huge AC, Adreani NM, Colombelli-Négrel D, Akçay C, Common LK, Kleindorfer S (2023). Age effects in Darwin’s finches: older males build more concealed nests in areas with more heterospecific singing neighbours. Journal of Ornithology.

Katsis, A.C., A.T.D. Bennett, K.L. Buchanan, S. Kleindorfer & M.M. Mariette (2023) Prenatal sound experience affects song preferences in male zebra finches. Animal Behaviour199, 1-9. doi:

Katsis, A. C., Evans, C., Common, L. K., Colombelli-Négrel D., & Kleindorfer S. (2023). Intraspecific fledgling adoption by a pair of Superb Fairy-wrens Malurus cyaneus. Australian Field Ornithology, 40, 49-55.

Katsis, A., Common, L., Hauber, M., Colombelli-Négrel, D., & Kleindorfer, S. (2023). From nestling to adult: personality traits are consistent within but not across life stages in a wild songbird, Behaviour (published online ahead of print 2023). doi:

Kleindorfer S, Heger B, Tohl D, Frigerio D, Hemetsberger J, Fusani L, Fitch WT, Colombelli-Négrel D (2023). Cues to individuality in greylag goose faces: algorithmic discrimination and behavioral field tests. Journal of Ornithology

Lawson S L, Fernandez-Duque F, Enos JK, Kleindorfer S, Ward MP, Gill SA, Hauber ME (2023).  Absence of referential alarm calls in long-term allopatry from the referent: A casy study with Galapagos Yellow Warblers. Behavioral Ecology and Sociobiology 77:99

McClure, C. J. W., Vargas, F. H., Amar, A., Concepcion, C. B., MacColl, C., & Sumasgutner, P. (2023). Conservation Letter: Monitoring Raptor Populations – A Call for Increased Global Collaboration and Survey Standardization. Journal of Raptor Research, 57(1), 1-8.

Nebel C, Haring E, Murgatroyd M, Sumasgutner SC, Gombobaatar S, Sumasgutner P, Zachos FE: Genetic Analysis of Golden Eagles (Aquila chrysaetos) from the Mongol-Altai: A Hotspot of Diversity and Implications for Global Phylogeography. J Raptor Res 2023, 57(3):1-16, 16.

Nwaogu, C. J., Amar, A., Nebel, C., Isaksson, C., Hegemann, A., & Sumasgutner, P. (2023). Innate immune function and antioxidant capacity of nestlings of an African raptor covary with the level of urbanisation around breeding territories. Journal of Animal Ecology, 92(1), 124-141.

Puehringer-Sturmayr V, Fiby M, Bachmann S, Filz S, Grassmann I, Hoi T, Janiczek C, Frigerio D. 2023. Effects of food-based enrichment on enclosure use and behavioral patterns in captive mammalian predators: a case study from an Austrian wildlife park.

Puehringer-Sturmayr, V., Krejci, J., Schuster, R., Kleindorfer, S., Kotrschal, K., Frigerio, D., & Loretto, M. (2023). Space use and site fidelity in the endangered Northern Bald Ibis Geronticus eremita: Effects of age, season, and sex. Bird Conservation International, 33(1), [E10].

Pühringer-Sturmayr V, Rittenschober J, Gegendorfer G, Kleindorfer S, Frigerio D (2023). Assessing quality of contributions to avian monitoring by non-scientists: a case study on individually banded wild birds. Environ. Res. Lett. 18.

Rittenschober J., Kleindorfer S., Frigerio D. The project design influences the quality of contributions in an online Citizen Science project, In Austrian Citizen Science Conference 2023 (ACSC2023) [014] Sissa Medialab. Proceedings of Science (PoS) Vol. 442 . DOI:

Sumasgutner, P., Cunningham, S. J., Hegemann, A., Amar, A., Watson, H., Nilsson, J. F., Andersson, M. N., & Isaksson, C. (2023). Interactive effects of rising temperatures and urbanisation on birds across different climate zones: A mechanistic perspective. Global Change Biology.

Sumasgutner, P., Nilles, T., Hykollari, A., de Chapa, M.M., Isaksson, C., Hochleitner, L., Renner, S. & Fusani, L. (2023) Integument colouration and circulating carotenoids in relation to urbanisation in Eurasian kestrels (Falco tinnunculus). The Science of Nature, 110.



Adreani, N. M., Valcu, M., Citizen Scientists, & Mentesana, L. (2022). Asymmetric architecture is non-random and repeatable in a bird's nest. Current Biology, 32(9), R412-413.

Bilby, J., Colombelli-Négrel, D., Katsis, A. C. & Kleindorfer, S. (2022). When aggressiveness could be too risky: linking personality traits and predator response in superb fairy-wrens. PeerJ10, [e14011].

Boucherie, P. H., Gallego-Abenza, M., Massen, J. J. & Bugnyar, T. (2022). Dominance in a socially dynamic setting: hierarchical structure and conflict dynamics in ravens' foraging groups. Philosophical Transactions of the Royal Society B377(1845), [20200446].

Colombelli-Négrel, D., Katsis, A. C., & Kleindorfer, S. (2022). Superb fairy-wrens with extreme exploration phenotypes respond more strongly to simulated territory intrusions. Animal Behaviour, 193, 101-111.

Common, L. K., Sumasgutner, P, Sumasgutner, S. C., Colombelli-Négrel, D., Dudaniec, R. Y., & Kleindorfer, S. (2022). Temporal and spatial variation in sex-specific abundance of the avian vampire fly (Philornis downsi). Parasitology Research, 121(1), 63-74.

Gallego-Abenza, M., Boucherie, P. H. & Bugnyar, T. (2022). Early social environment affects attention to social cues in juvenile common ravens, Corvus corax. Royal Society Open Science9(6), [220132].

Gallup, A. C., Schild, A. B., Ühlein, M. A., Bugnyar, T. & Massen, J. J. (2022). No Evidence for Contagious Yawning in Juvenile Ravens (Corvus corax): An Observational Study. Animals12(11), 1357.

García Loor, J., Espinosa, R., Alarcón, I., Luzuriaga, N., & Guevara, E. (2022). Notas sobre la dieta del Zambullidor Plateado Norteño Podiceps occipitalis juninensis. Revista Ecuatoriana De Ornitología8(2), 71-73.

García-Loor, J. (2022). Elevational range extension of Drab Water Tyrant, Ochthornis littoralis (Pelzeln, 1868) (Passeriformes, Tyrannidae). Check List, 18(3), 451-453.

Gegendorfer, G., Rittenschober, J., Vesely, H. & Frigerio D. (2022). Volunteering In Monitoring Bird Behavior: Accuracy Matters! Frontiers for Young Minds, 10, [596181].

Guggenberger, M., Adreani, N. M., Foerster, K., & Kleindorfer, S., (2022). Vocal recognition of distance calls in a group-living basal bird: the greylag goose, Anser anser. Animal behaviour. 186, 107-119.

Hirschenhauser, K., Frigerio, D., & Neuböck-Hubinger, B. (2022). Science Education and Beyond: Citizen Science in Primary School Potentially Affects Conceptual Learning and Socio-emotional Development. In A. Volvlas (Hrsg.), Citizen Science - Methods, Approaches and New Perspectives [Working Title]. Intechopen.

Horn, L., Cimarelli, G., Boucherie, P. H., Šlipogor, V. & Bugnyar, T. (2022). Beyond the dichotomy between field and lab—the importance of studying cognition in context. Current Opinion in Behavioral Sciences46, [101172].

Jain, V., Bugnyar, T., Cunningham, S. J., Gallego-Abenza, M., Loretto, M.-C., & Sumasgutner, P. (2022). The spatial and temporal exploitation of anthropogenic food sources by common ravens (Corvus corax) in the Alps. Movement Ecology, 10, [35].

Kleindorfer, S., Colombelli-Négrel, D., Common, L. K., O'Connor, J. A., Peters, K. J., Katsis, A. C., Dudaniec, R. Y., Sulloway, F. J., & Adreani, N. M. (2022). Functional traits and foraging behaviour: Avian vampire fly larvae change the beak and fitness of their Darwin's finch hosts. Functional Ecology, 36(7), 1806–1817.

Kotrschal, K. (2022) Warum wir einander verstehen und gut zusammenleben könn(t)en. in: Jessica Ullrich (ed.): Kohabitation, Koexistenz, Konvivialität. Tierstudien 22, Neofelis Verlag

Kotrschal, K. (2022) Zivilgesellschaft, liberale Demokratie und der Schutz der Umwelt. in: P. Hilpold, W. Steinmair & A. Raffeiner (Hg.): Österreich und die EU im Umbruch – eine Nachlese zur Festschrift für Heinrich Neisser (p 202-210). Facultas

Meixner, J., & Kotrschal, K. (2022). Animal-Assisted Interventions With Dogs in Special Education—A Systematic Review. Frontiers in Psychology, 13, [876290].

Miller, R., Lambert, M. L., Frohnwieser, A., Brecht, K. F., Bugnyar, T., Crampton, I., Garcia-Pelegrin, E., Gould, K., Greggor, A. L., Izawa, E.-I., Kelly, D. M., Li, Z., Luo, Y., Luong, L. B., Massen, J. J. M., Nieder, A., Reber, S. A., Schiestl, M., Seguchi, A., Sepehri, P., Stevens, J. R., Taylor, A. H., Wang, L., Wolff, L. M., Zhang, Y. & Clayton, N. S. (2022). Socio-ecological correlates of neophobia in corvids. Current Biology32(1), 74-85.

Müller, K., Lengheimer, T., Kral-Pointner, J. B., Wojta, J., Yeghiazaryan, L., Krall, C., Palme, R., Kleindorfer, S., Plasenzotti, R., Pollak, D. D. & Tillmann, K. E. (2022). Exposure to soiled bedding reduces abnormal repetitive behaviors in mice. Frontiers in Behavioral Neuroscience, 16, [1062864].

Naikatini, A. N., Keppel, G., Brodie, G., & Kleindorfer, S. (2022). Interspecific Competition and Vertical Niche Partitioning in Fiji’s Forest Birds. Diversity, 14(3), [223].

Naikatini, A. N., Keppel, G., Brodie, G., & Kleindorfer, S. (2022). Avian diversity and abundance across years: consistent patterns in forests but not grasslands on Viti Levu, Fiji. Pacific Conservation Biology

Stofberg, M., Amar, A., Sumasgutner, P., & Cunningham, S. J. (2022). Staying cool and eating junk: Influence of heat dissipation and anthropogenic food on foraging and body condition in an urban passerine. Landscape and Urban Planning, 226, [104465].

Wascher, C. A. F., Arnold, W., & Kotrschal, K. (2022). Effects of severe anthropogenic disturbance on the heart rate and body temperature in free-living greylag geese (Anser anser). Conservation Physiology, 10(1), [coac050].



Antonson, N. D., Rivera, M., Abolins-Abols, M., Kleindorfer, S., Liu, W.-C., & Hauber, M. E. (2021). Early auditory experience alters genome-wide methylation in the auditory forebrain of songbird embryos. Neuroscience Letters, 755, [135917].

Bates, A. E., Primack, R. B., PAN-Environment Working Group, Duarte C. M. (2021). Global COVID-19 lockdown highlights humans as both threats and custodians of the environment. Biological Conservation, 263, [109175].

Böhm, C., Bowden, C. G. R., Seddon, P. J., Hatipoglu, T., Oubrou, W., El Bekkay, M., Quevedo, M. A., Fritz, J., Yeniyurt, C., Lopez, J. M., Orueta, J. F., Frigerio, D., & Unsöld, M. (2021). The northern bald ibis Geronticus eremita: history, current status and future perspectives. Oryx, 55(6), 934-946.

Catto, S., Sumasgutner, P., Amar, A., Thomson, R. L., & Cunningham, S. J. (2021). Pulses of anthropogenic food availability appear to benefit parents, but compromise nestling growth in urban Red-winged Starlings. Oecologia, 197(3), 565-576.

Colombelli-Négrel, D., & Kleindorfer, S. (2021). Behavioural response to songs between genetically diverged allopatric populations of Darwin's small tree finch in the Galápagos. Journal of Evolutionary Biology, 34(5), 816-829.

Colombelli-Négrel, D., Hauber, M. E., Evans, C., Katsis, A. C., Brouwer, L., Adreani, N. M., & Kleindorfer, S. (2021). Prenatal auditory learning in avian vocal learners and non-learners. Philosophical Transactions of the Royal Society B, 376(1836), [20200247].

Common, L. K., Sumasgutner, P., Dudaniec, R. Y., Colombelli-Négrel, D., & Kleindorfer, S. (2021). Avian vampire fly (Philornis downsi) mortality differs across Darwin’s finch host species. Scientific Reports, 11, [15832].

Ersoy, S., Maag, N., Boehly, T., Boucherie, P. H. & Bugnyar, T. (2021). Sex-specific parental care during postfledging in common ravens. Animal Behaviour, 181, 95-103.

Frigerio, D., Richter, A., Per, E., Pruse, B., & Vohland, K. (2021). Citizen Science in natural sciences. In: K. Vohland, A. Land, L. Ceccaroni, R. Lemmens, J. Perelló, M. Ponti, R. Samson, & K. Wagenknecht (eds), The Science of Citizen Science (Chapter 5). Springer.

Frigerio, D., Sumasgutner, P., Kotrschal, K., Kleindorfer, S., & Hemetsberger, S. (2021). From individual to population level: Temperature and snow cover modulates fledging success through breeding phenology in Greylag geese (Anser anser). Scientific Reports, 11, [16100].

Gallego-Abenza, M., Blum, C., & Bugnyar, T. (2021). Who is crying wolf? Seasonal effect on anti-predator response to age-specific alarm-calls in common ravens, Corvus corax. Learning & Behavior. 49(1), 159-167.

Katsis, A. C., Buchanan, K. L., Kleindorfer, S., & Marriette, M. M. (2021). Long-term effects of prenatal sound experience on songbird behavior and their relation to song learning. Behavioral Ecology and Sociobiology, 75(1), 18.

Katsis, A. C., Colombelli-Négrel, D., Common, L. K., O’Connor, J. A., Dudaniec, R. Y., Garcia-Loor, J., & Kleindorfer, S. (2021). Nestling behaviour predicts naris deformation in Darwin’s finches parasitised parasitized by the avian vampire fly. Biological Journal of the Linnaean Society, 134(3), 636-649.

Kislinger, L., & Kotrschal, K. (2021). Hunters and Gatherers of Pictures: Why Photography Has Become a Human Universal. Frontiers in Psychology, 12, [654474].

Kleindorfer, S., Common, L. K., & Sumasgutner, P. (2021). Nesting Success and Nesting Height in the Critically Endangered Medium Tree Finch (Camarhynchus pauper). Birds, 2(4), 427-444.

Kleindorfer, S., Common, L. K., O’Connor, J. A., García Loor, H. J., Katsis, A. C., Dudaniec, R. Y., Colombelli-Négrel, D., & Adreani, M. N. (2021). Female in-nest attendance predicts the number of ectoparasites in Darwin’s finch species. Proceedings of the Royal Society of London B: Biological Sciences, 288(1965), [20211668].

Kotrschal, K. (2021). Die Natur des Menschen und die anderen Tiere: Tierschutz in evolutionärer Perspektive. In S. Horstmann (Ed.), Interspezies Lernen: Grundlinien interdisziplinärer Tierschutz- und Tierrechtsbildung (pp. 61-88). Transcript Verlag. Human-animal studies Vol. 27

McCarren, S., Sumasgutner, P., Tate, G., Koeslag, A., & Amar, A. (2021). Clinal variation in the polymorphic Black Sparrowhawk Accipiter melanoleucus is unrelated to infection by the blood parasite Haemoproteus nisi. Journal of Ornithology, 162(1), 231-241.

McPherson, S. C., Sumasgutner, P., & Downs, C. T. (2021). South African raptors in urban landscape: a review. Ostrich, 92(1), 41-57.

Mentesana, L., & Adreani, N. M. (2021). Acute aggressive behavior perturbates the oxidative status of a wild bird independently of testosterone and progesterone. Hormones and Behavior, 128, [104913].

Nebel, C., Amar, A., Hegemann, A., Isaksson C., & Sumasgutner, P. (2021). Parental morph combination does not influence innate immune function in nestlings of a colour-polymorphic African raptor. Scientific Reports, 11, [11053].

Nebel, C., Sumasgutner, P., Rodseth, E., Ingle, R. A., Childs, D. Z., Curtis-Scott, O., & Amar, A. (2021). Multigenerational pedigree of wild individually marked black sparrowhawks suggests that dark plumage colouration is dominantly inherited. Journal of Zoology, 315(2), 123-130.

Neuböck-Hubinger, B., Grubner, K., Frigerio, D., & Hirschenhauser, K. (2021). Natural science education – a claim for equality, diversity and inclusion in Citizen Science activities. In Austrian Citizen Science Conference 2020 (ACSC2020) [018] Sissa Medialab. Proceedings of Science (PoS) Vol. 393

Odom, K. J., Cain, K. E., Hall, M. L., Langmore, N. E., Mulder, R. A., Kleindorfer, S., Karubian, J., Brouwer, L., Enbody, E. D., Jones, J. A. and, Dowling, J. L., Leitao, A. V., Graig, E. I., Evans, C., Johnson, A. E., Meyers, K. K-A., Araya Salas, M., & Webster, M. S. (2021). Sex role similarity and sexual selection predict male and female song elaboration and dimorphism in fairy‐wrens. Ecology and Evolution, 11(24), 17901-17919.

Puehringer-Sturmayr, V., Stiefel, T., Kotrschal, K., Kleindorfer, S., & Frigerio, D. (2021). Social interactions change with season and age in Northern Bald Ibis. Journal of Ornithology, 162(1), 277–288.

Reynolds, C., Byrne, M. J., Chamberlain, D. E., Howes, C. G., Seymour, C. L., Sumasgutner, P., & Taylor, P. J. (2021). Urban Animal Diversity in the Global South. In: C. M. Shackleton, S. S. Cilliers, E. Davoren, M. J. du Toit (Eds), Urban Ecology in the Global South (p. 169-202). Springer International Publishing.

Risi, T. C., Sumasgutner, P., & Cunningam, S. J. (2021).  Anthropogenic food availability and body mass maintenance in urban Red-winged Starlings Onychognathus morio. Ostrich, 92(1), 16-25.

Rittenschober, J., Pühringer-Sturmayr, V., & Frigerio, D. (2021). The relevance of a digital platform for the citizen science project NestCams. In Austrian Citizen Science Conference 2020 (ACSC2020) [009] Sissa Medialab. Proceedings of Science (PoS) Vol. 393

Stocker, M., Prosl, J., Vanhooland, L. C., Horn, L., Bugnyar, T., Canoine, V. & Massen, J. J. (2021). Measuring salivary mesotocin in birds-Seasonal differences in ravens' peripheral mesotocin levels. Hormones and Behavior, 134, [105015].

Sumasgutner, P. (2021). Guest Editorial: A Southern perspective on urban birds. Ostrich: Journal of African Ornithology, 92(1), III–V.

Sumasgutner, P., Buij, R., McClure, C. J. W., Shaw, P., Dykstra, C. R., Kumar, N., & Rutz, C. (2021). Raptor research during the COVID-19 pandemic provides invaluable opportunities for conservation biology. Biological Conservation, 260, [109149].

Wascher, C. A. F., Allen, K., & Szipl, G., (2021). Learning and motor inhibitory control in crows and domestic chickens. Royal Society Open Science, 8(10), [210504].

Wascher, C. A. F., Baur, N., Hengl, M., Köck, C., Pegger, T., Schindlbauer, J., & Wemer, L. (2021). Behavioral responses of captive corvids to the presence of visitors. Animal Behavior and Cognition, 8(4), 481-492.

Wemer, L., Hegemann, A., Isaksson, C., Nebel, CarinaC., Kleindorfer, S., Gamauf, A., Adrion, M., & Sumasgutner, P. (2021). Reduced ectoparasite load, body mass and blood haemolysis in Eurasian kestrels (Falco tinnunculus) along an urban-rural gradient. The Science of Nature, 108(5), [42].

Wenig, K., Boucherie, P. H., & Bugnyar, T. (2021). Early evidence for emotional play contagion in juvenile ravens. Animal Cognition, 24(4), 717-729, 1-13.

Westreicher, F., Cieslinski, M., Ernst, M., Frigerio, D., Heinisch, B., Hübner, T., & Rüdisser, J. (2021). Recognizing failures in citizen science projects: Lessons learned. In Austrian Citizen Science Conference 2020 (ACSC2020) [007] Sissa Medialab. Proceedings of Science (PoS) Vol. 393



Adreani, N. M., D'Amelio, P. B., Gahr, M., & ter Maat, A. (2020). Life-Stage Dependent Plasticity in the Auditory System of a Songbird is Signal and Emitter-Specific. Frontiers in Neuroscience, 14, [588672].

Beck, K., Loretto, M.-C., & Bugnyar, T. (2020). Effects of site fidelity, group size and age on food caching behaviour of common ravens, Corvus corax. Animal Behaviour, 164, 51-64.

Common, L. K., Dudaniec, R. Y., Colombelli-Négrel, D., & Kleindorfer, S. (2020). Taxonomic shifts in Philornis larval behaviour and rapid changes in Philornis downsi Dodge & Aitken (Diptera: Muscidae): an invasive avian parasite on the Galápagos Islands. In M. Sarwar (eds), Life Cycle and Development of Diptera. Intechopen.

Common, L. K., O’Connor, J. A., Dudaniec, R. Y., Peters, K. J., & Kleindorfer, S. (2020). Evidence for rapid downward fecundity selection in an ectoparasite (Philornis downsi) with earlier host mortality in Darwin’s finches. Journal of Evolutionary Biology, 33(4), 524-533.

Davidkova, M., Vesely, P., Syrova, M., Nacarova, J., & Bugnyar, T. (2020). Raven respond to unfamiliar corvid alarm calls. Journal of Ornithology, 161(4), 967-975.

Frigerio, D., Pühringer-Sturmayr, V., Steinbacher, C., Rittenschober, J., & Kleindorfer, S. (2020). Citizen science, animal behaviour and digital devices: challenges and solutions. Paper presented at 5th Austrian Citizen Science Conference 2019 (ACSC2019), Obergurgl, Austria.

Gallego-Abenza, M., Loretto, M.-C., & Bugnyar, T. (2020). Decision time modulates social foraging success in wild common ravens, Corvus corax. Ethology, 126(4), 413-422.

Gallego-Abenza, M., Mathevon, N., & Wheatcroft, D. (2020). Experience modulates an insect's response to anthropogenic noise. Behavioral Ecology, 31(1), 90-96.

Gegendorfer, G., & Frigerio, D. (2020). How having a family improves digestion in social geese. Frontiers for Young Minds, 8, [540008].

Heigl, F., Kieslinger, B., Paul, K. T., Uhlik, J., Frigerio, D., & Dörler, D. (2020). Co-Creating and Implementing Quality Criteria for Citizen Science. Citizen Science: Theory and Practice, 5(1): 23, 1-11.

Huchler, K., Schulze, C. H., Gamauf, A., & Sumasgutner, P. (2020). Shifting Breeding Phenology in Eurasian Kestrels Falco tinnunculus: Effects of Weather and Urbanization. Frontiers in Ecology and Evolution, 8, [247].

Kleindorfer, S., & Dudaniec, R. Y. (2020). Hybridization fluctuates with rainfall in Darwin's tree finches. Biological Journal of the Linnaean Society, 130(1), 79-88.

Loretto, M.-C., Schuster, R., Federspiel, I. G., Heinrich, B., & Bugnyar, T. (2020). Contextual imitation in juvenile common ravens (Corvus corax). Animal Behaviour, 163, 127-134.

Mentesana, L., Moiron, M., Guedes, E., Cavalli, E., Tassino, B., & Adreani, N. M. (2020). Defending as a unit: sex- and context-specific territorial defence in a duetting bird. Behavioral Ecology and Sociobiology, 74(9), [111].

Mueller, R., Amar, A., Sumasgutner, P., McPherson, S. C., & Downs, C. (2020). Urbanization is associated with increased breeding rate, but decreased breeding success, in an urban population of near-threatened African Crowned Eagles. The Condor, 122(3), 1-11.

Nebel, C., Harl, J., Pajot, A., Weissenböck, H., Amar, A., & Sumasgutner, P. (2020). High prevalence and genetic diversity of Haemoproteus columbae (Haemosporida: Haemoproteidae) in feral pigeons Columba livia in Cape Town, South Africa. Parasitology Research, 119(2), 447-463.

Nebel, C., Sumasgutner, P., McPherson, S. C., Tate, G. J., & Amar, A. (2020). Contrasting parental color morphs increase regularity of prey deliveries in an African raptor. Behavioral Ecology, 31(5), 1142–1149.

Puehringer-Sturmayr, V., Hemetsberger, F., & Frigerio, D. (2020). Tracking Birds: How a Little Backpack Could Affect the Life of a Wild Bird. Frontiers for Young Minds, 8, [116].

Puehringer-Sturmayr, V., Loretto, M. C., Hemetsberger, J., Czerny, T., Gschwandegger, J., Leitsberger, M., Kotrschal, K., & Frigerio, D. (2020). Effects of bio-loggers on behaviour and corticosterone metabolites of Northern Bald Ibises (Geronticus eremita) in the field and in captivity. Animal Biotelemetry, 8, [2].

Sierro, J., Loretto, M.-C., Szipl, G., Massen, J. J. M., & Bugnyar, T. (2020) Food calling in ravens (Corvus corax) revisited: who is addressed? Ethology, 126(2), 257-266.

Sumasgutner, P., Jenkins, A., Amar, A., & Altwegg, R. (2020). Nest boxes buffer the effects of climate on breeding performance in an African urban raptor. PLoS ONE, 15(6), [e0234503].



Frigerio, D., Hemetsberger, F., & Wascher, C. A. F. (2019). How having a partner and a family changes the stress levels of greylag geese. Frontiers for Young Minds, 7, [68].

Frigerio, D., Puehringer-Sturmayr, V., Neuböck-Hubinger, B., Gegendorfer, G., Kotrschal, K., & Hirschenhauser, K. (2019). Monitoring public awareness about the endangered northern bald ibis: a case study involving primary school children as citizen scientists. PeerJ, 7, [e7569].

Hebesberger, D. V., Beetz, A., & Kotrschal, K. (2019). Behavioral and Physiological Effects of Equine-assisted Early Intervention for Mother- Child Dyads with Insecure Attachment. Human-Animal Interaction Bulletin, 24-37.

Hirschenhauser, K., Frigerio, D., Leighinger, V., Schenkenfelder, I., & Neuböck-Hubinger, B. (2019). Primary pupils, science and a model bird species: evidence for the efficacy of extracurricular science education. PLoS ONE, 14(7), [e0220635].

Kleindorfer, S., Custance, G., Peters, K. J., & Sulloway, F. J. (2019). Introduced parasite changes host phenotype, mating signal and hybridization risk: Philornis downsi effects on Darwin's finch song. Proceedings of the Royal Society B: Biological Sciences, 286(1904), [20190461].

Kortekaas, K., & Kotrschal, K. (2019). Does socio-ecology drive differences in alertness between wolves and dogs when resting? Behavioural Processes, 166, [103877].

Langton, A., & Kleindorfer, S. (2019). Minimum longevity and age-related male plumage in Darwin's finches on Floreana Island. Journal of Ornithology, 160(2), 351-361.

Lesch, R., Kotrschal, K., Schöberl, G. I., Beetz, A., Solomon, J., & Fitch, W. T. (2019). Talking to Dogs: Companion Animal-Directed Speech in a Stress Test. Animals, 9(7), [417].

Loo, W. T., Dudaniec, R. Y., Kleindorfer, S., & Cavanaugh, C. M. (2019). An inter-island comparison of Darwin’s finches reveals the impact of habitat, host phylogeny, and island on the gut microbiome. PLoS ONE, 14(12), [e0226432].

Loo, W. T., Dudaniec, R. Y., Loor, J. G., Kleindorfer, S., & Cavanaugh, C. M. (2019). Host phylogeny, diet, and habitat differentiate the gut microbiomes of Darwin’s finches on Santa Cruz Island. Scientific Reports, 9, [18781].

Peters, K. J., Evans, C., Aguirre, J. D., & Kleindorfer, S. (2019). Genetic admixture predicts parasite intensity: evidence for increased hybrid performance in Darwin's tree finches. Royal Society Open Science, 6(4), [181616].

Richter, A., Sieber, A., Siebert, J., Miczajka-Rußmann, V. L., Zabel, J., Ziegler, D., Hecker, S., & Frigerio, D. (2019). Storytelling for narrative approaches in citizen science: towards a generalized model. Journal of Science Communication, 18(06), [A02].

Rivera, M., Cealie, M., Hauber, M. E., Kleindorfer, S., & Liu, W. (2019). Neural activation in response to conspecific songs in zebra finch (Taeniopygia guttata) embryos and nestlings. NeuroReport, 30(3), 217-221.

Solomon, J., Beetz, A., Schöberl, G. I., Gee, N., & Kotrschal, K. (2019). Attachment security in companion dogs: adaptation of Ainsworth’s strange situation and classification procedures to dogs and their human caregivers. Attachment and Human Development, 21(4), 389-417.

Szipl, G., Depenau, M., Kotrschal, K., Hemetsberger, J., & Frigerio, D. (2019). Costs and benefits of social connectivity in juvenile greylag geese. Scientific Reports, 9, [12839].

Szipl, G., Loth, A., Wascher, C.A.F., Hemetsberger, J., Kotrschal, K., & Frigerio, D. (2019). Parental behaviour and family proximity as key to gosling survival in Greylag Geese (Anser anser). Journal of Ornithology, 160(2), 473-483.



Boeckle, M., Szipl, G., & Bugnyar, T. (2018). Raven food calls indicate sender's age and sex. Frontiers in Zoology, 15, [5].

Frigerio, D., Kotrschal, K., Fabro, C., Puehringer-Sturmayr, V., Iaiza, L., Hemetsberger, J., Mason, F., Sarnataro, C., & Filacorda, S. (2018). Social context modulates digestive efficiency in greylag geese (Anser anser). Scientific reports, 8, [16498].

Frigerio, D., Pipek, P., Kimmig, S., Winter, S., Melzheimer, J., Diblíková, L., Wachter, B., & Richter, A. (2018). Citizen science and wildlife biology: Synergies and challenges. Ethology, 124(6), 365-377.

Puehringer-Sturmayr, V., Gegendorfer, G., Rittenschober, J., Szipl, G., Frigerio, D. (2018). Involving pupils/citizens in long-term behavioural biology research: Lessons learnt and future perspectives. In F. Heigl, D. Dörler, M. Ernst, (eds), Proceedings of the Austrian Citizen Science Conference, 2018 Februray 1-3, Salzburg, Austria. (p. 34-38.). Frontiers.

Puehringer-Sturmayr, V., Wascher, C. A. F., Loretto, M.-C., Palme, R., Stoewe, M., Kotrschal, K., & Frigerio, D. (2018). Seasonal differences of corticosterone metabolite concentrations and parasite burden in Northern Bald Ibis (Geronticus eremita): The role of affiliative interactions. PLoS ONE, 13(1), [e0191441].

Ringler, E., Szipl, G., Harrigan, R. J., Bartl-Binder, P., Mangione, R., & Ringler, M. (2018). Hierarchical decision‐making balances current and future reproductive success. Molecular Ecology, 27(9), 2289–2301.

Szipl, G., Ringler, E., & Bugnyar, T. (2018). Attacked ravens flexibly adjust signalling behaviour according to audience composition. Proceedings of the Royal Society of London Series B: Biological Sciences, 285(1880), [20180375].

Wascher, C. A. F., Kotrschal, K., & Arnold, W. (2018). Free-living greylag geese adjust their heart rates and body core temperatures to season and reproductive context. Scientific Reports8, [2142].

Wirtz, S., Böhm, C., Fritz, J., Kotrschal, K., Veith, M., & Hochkirch, A. (2018). Optimizing the genetic management of reintroduction projects: genetic population structure of the captive Northern Bald Ibis population. Conservation Genetics, 19(4), 853–864.



Frigerio, D., Ludwig, S. C., Hemetsberger, J., Kotrschal, K., & Wascher, C. A. F. (2017). Social and environmental factors modulate leucocyte profiles in free-living Greylag geese (Anser anser). PeerJ5, [e2792].

Kotrschal, A., Zeng, H-L., van der Bijl, W., Öhman-Mägi, C., Kotrschal, K., Pelckmans, K., & Kolm, N. (2017). Evolution of brain region volumes during artificial selection for relative brain size. Evolution, 71(12), 2942-2951.

Kotrschal, K. (2017). Konrad Lorenz. In J. Vonk, & T. K. Shackelford (Eds.), Encyclopedia of Animal Cognition and Behavior (1 ed.). Springer International Publishing.

Kotrschal, K. (2017). Vom wilden Wolf zum treuen Freund. In S. Florian (Ed.), Unsere Hunde Herr am Hof Verlag.

Kotrschal, K. (2017). Warum wirken Tiere in der Pädagogik? Erziehung und Unterricht, 9-10, 696-706.

Loretto, M.-C., Schuster, R., Itty, C., Marchand, P., Genero, F., & Bugnyar, T. (2017). Fission-fusion dynamics over large distances in raven non-breeders. Scientific Reports7, [380].

Loth, A., Frigerio, D., Kotrschal, K., & Szipl, G. (2017). Differential responses to gosling distress calls in parental and non-parental Greylag Geese. Journal of Ornithology, 159(2), 401-412.

Ludwig, S. C., Kapetanopoulos, K., Kotrschal, K., & Wascher, C. A. F. (2017). Effects of mate separation in female and social isolation in male free-living Greylag geese on behavioural and physiological measures. Behavioural Processes, 138, 134-141.

Schöberl, G. I., Wedl, M., Beetz, A., & Kotrschal, K. (2017). Psychobiological Factors Affecting Cortisol Variability in Human-Dog Dyads. PLoS ONE, 12(2), [e0170707].

Szipl, G., Ringler, E., Spreafico, M., & Bugnyar, T. (2017). Calls during agonistic interactions vary with arousal and raise audience attention in ravens. Frontiers in Zoology, 14, [57].

Wascher, C. A. F., Hemetsberger, J., Kotrschal, K., & Frigerio, D. (2017). Leucocyte profiles and family size in fledgling Geylag Geese (Anser anser). Avian Biology Research10(4), 246–252.



Beck, K., Frigerio, D., & Loretto, M.-C. (2016). Der Kolkrabe – Schwarzfedriges Genie oder doch nur ein Unglücksbote? - Von Irrtümern und Mythen bis hin zur Forschung von Groß und Klein. Öko-L38, 3–11.

Frigerio, D., Cibulski, L., Ludwig, S. C., Campderrich, I., Kotrschal, K., & Wascher, C. A. F. (2016). Excretion patterns of coccidian oocysts and nematode eggs during the reproductive season in Northern Bald Ibis (Geronticus eremita). Journal of Ornithology157(3), 839–851.

Hirschenhauser, K., Frigerio, D., & Neuböck-Hubinger, B. (2016). Wirkungen außerschulischer Angebote im Sachunterricht: das Waldrapp-Projekt - Impact of external projects on teaching natural sciences in primary school: the Northern bald ibis case. In Transfer Forschung-Schule “Visible Didactics - Fachdidaktische Forschung trifft Praxis” (pp. 307–308). Bad Heilbrunn: Linkhard Verlag.

Kotrschal, K. (2016). Verwaltungsaufblähung und Verschulung? In P. Kellermann, H. Guggenberger, & K. Weber (Eds.), Universität nach Bologna? Hochschulkonzeptionen zwischen Kritik und Utopie Mandelbaum Verlag.

Kotrschal, K., Scheiber, I. B. R., & Hirschenhauser, K. (2016). Individual Performance in Complex Social Systems: The Greylag Goose Example. In P. Kappeler (Ed.), Animal Behaviour: Evolution & Mechanisms (pp. 121-148). Heidelberg: Springer.

Loretto, M.-C., Schuster, R., & Bugnyar, T. (2016). GPS tracking of non-breeding ravens reveals the importance of anthropogenic food sources during their dispersal in the Eastern Alps. Current Zoology62(4), 337–344.

Schöberl, G. I., Beetz, A., Solomon, J., Wedl, M., Gee, N., & Kotrschal, K. (2016). Social factors influencing cortisol modulation in dogs during a strange situation procedure. Journal of Veterinary Behavior: Clinical Applications and Research, 11, 77-85.



Beetz, A., & Kotrschal, K. (2015). Was die Wissenschaft weiß – Ein Überblick über internationale Forschungsergebnisse im Bereich Mensch-Tier-Interaktion. In Heimtiere und Gesundheit. Prävention – Assistenz – Therapie (pp. 6-23). Mars Petcare Deutschland GmbH.

Kotrschal, K. (2015). Biologie oder Moral? In H. Barta, M. Lang, & R. Rollinger (Eds.), Prozeßrecht und Eid: Recht und Rechtsfindung in antiken Kulturen Philippika: altertumswissenschaftliche Abhandlungen Vol. 86

Loretto, M.-C., Reimann, S., Schuster, R., Graulich, D. M., & Bugnyar, T. (2015). Shared space, individually used: spatial behaviour of non-breeding ravens (Corvus corax) close to a permanent anthropogenic food source. Journal of Ornithology157(2), 439–450.

Schöberl, G. I., Kortekaas, K., Schöberl, F., & Kotrschal, K. (2015). Algorithm-supported visual error correction (AVEC) of heart rate measurements in dogs, Canis lupus familiaris. Behavior Research Methods, 47(4), 1356-1364.

Szipl, G., Boeckle, M., Wascher, C. A. F., Spreafico, M., & Bugnyar, T. (2015). With whom to dine? Ravens’ responses to food-associated calls depend on individual characteristics of the caller. Animal Behaviour99, 33–42.

Wedl, M., Kotrschal, K., Julius, H., & Beetz, A. (2015). Children with Avoidant or Disorganized Attachment Relate Differently to a Dog and to Humans During a Socially Stressful Situation. Anthrozoös: a multidisciplinary journal of the interactions of people and animals, 28(4), 601-610.



Cibulski, L., Wascher, C. A. F., Weiß, B. M., & Kotrschal, K. (2014). Familiarity with the experimenter influences the performance of Common ravens (Corvus corax) and Carrion crows (Corvus corone corone) in cognitive tasks. Behavioural Processes103, 129–137.

Dorn, S., Wascher, C. A. F., Möstl, E., & Kotrschal, K. (2014). Ambient temperature and air pressure modulate hormones and behaviour in Greylag geese (Anser anser) and Northern bald ibis (Geronticus eremita). Behavioural Processes108, 27–35.

Frigerio, D., Hemetsberger, J. & Kotrschal, K. (2014). Selbst nach 40 Jahre Forschung: Neues von den Grünauer Graugänsen - Von internationaler Grundlagenforschung bis zur Zusammenarbeit mit Volksschulkindern. Öko-L36, 3–13.

Hillemann, F., Bugnyar, T., Kotrschal, K., & Wascher, C. A. F. (2014). Waiting for better, not for more: corvids respond to quality in two delay maintenance tasks. Animal Behaviour90, 1–10.

Massen, J. J. M., Pašukonis, A., Schmidt, J., & Bugnyar, T. (2014). Ravens notice dominance reversals among conspecifics within and outside their social group. Nature Communications5, [3679].

Massen, J. J. M., Szipl, G., Spreafico, M., & Bugnyar, T. (2014). Ravens intervene in others’ bonding attempts. Current Biology24(22), 2733–2736.

Szipl, G., & Bugnyar, T. (2014). Craving ravens: Individual “haa” call rates at feeding sites as cues to personality and levels of fission-fusion dynamics? Animal Behavior and Cognition1(3), 265–280.

Szipl, G., Boeckle, M., Werner, S. A. B., & Kotrschal, K. (2014). Mate Recognition and Expression of Affective State in Croop Calls of Northern Bald Ibis (Geronticus eremita). PLoS ONE9(2). [e88265].

Ujfalussy, D. J., Miklósi, Á., Bugnyar, T., & Kotrschal, K. (2014). Role of mental representations in quantity judgments by jackdaws (Corvus monedula). Journal of Comparative Psychology128(1), 11–20.



Bugnyar, T. (2013). Social cognition in ravens. Comparative Cognition & Behavior Reviews8, 1–12.

Frigerio, D., & Gegendorfer, G. (2013). Exotisch oder heimisch? - Der Waldrapp aus dem Almtal. Groß und Klein erforschen die vom Aussterben bedrohten Vogelarten. Öko-L35(3), 3–13.

Guibert, F., Lumineau, S., Kotrschal, K., Möstl, E., Richard-Yris, M.-A., & Houdelier, C. (2013). Trans-generational effects of prenatal stress in quail. Proceedings. Biological Sciences280(1753), [20122368].

Hirschenhauser, K., Gahr, M., & Goymann, W. (2013). Winning and losing in public: Audiences direct future success in Japanese quail. Hormones and Behavior63(4), 625–633.

Kotrschal, K. (2013). The Quest for Understanding Social Complexity. Quo Vadis, Behavioural Biology? Past, Present, and Future of an Evolving Science111(380), 77–88.

Mikolasch, S., Kotrschal, K., & Schloegl, C. (2013). Transitive inference in jackdaws. Behavioural Processes92, 113–117.

Schloegl, C., Waldmann, M. R., & Fischer, J. (2013). Understanding of and reasoning about object–object relationships in long-tailed macaques? Animal Cognition16(3), 493–507.

Teschke, I., Wascher, C. A. F., Scriba, M. F., von Bayern, A. M. P., Huml, V., Siemers, B., & Tebbich, S. (2013). Did tool-use evolve with enhanced physical cognitive abilities? Philosophical Transactions of the Royal Society of London. B, Biological Sciences368(1630), [20120418].

Wascher, C. A. F., & Bugnyar, T. (2013). Behavioral responses to inequity in reward distribution and working effort in crows and ravens. PLoS ONE8(2), [e56885].

Weiß, B. M., & Scheiber, I. B. R. (2013). Long-term memory of hierarchical relationships in free-living greylag geese. Animal Cognition16(1), 91–97.



Bisenberger, A., Frigerio, D. (2012). Biene Maja zeigt uns ihre Welt - von einer Projektidee zur angewandten Methodik tiergestützter Vermittlungsarbeit mit Kleintieren. Öko-L2, 29–35.

Boeckle, M., & Bugnyar, T. (2012). Long-term memory for affiliates in ravens. Current Biology22(9), 801–806.

Boeckle, M., Szipl, G., & Bugnyar, T. (2012). Who wants food? Individual characteristics in raven yells. Animal Behaviour84(5), 1123–1130.

Braun, A., & Bugnyar, T. (2012). Social bonds and rank acquisition in raven nonbreeder aggregations. Animal Behaviour84(6), 1507–1515.

Braun, A., Walsdorff, T., Fraser, O. N., & Bugnyar, T. (2012). Socialized sub-groups in a temporary stable Raven flock? Journal of Ornithology153(1), 97–104.

Frigerio, D., Kotrschal, K., Millesi, E., & Hemetsberger, J. (2012). Children and Scientific Observations: Pupils Measuring Greylag Goose Behaviour. International Journal for Cross-Disciplinary Subjects in Education, 3(4), 867–872.

Dufour, V., Wascher, C. A. F., Braun, A., Miller, R., & Bugnyar, T. (2012). Corvids can decide if a future exchange is worth waiting for. Biology Letters8(2), 201–204.

Fraser, O. N., & Bugnyar, T. (2012). Reciprocity of agonistic support in ravens. Animal Behaviour83(1), 171–177.

Hirschenhauser, K., Spreitzer, K., Lepschy, M., Kotrschal, K., & Möstl, E. (2012). Excreted corticosterone metabolites differ between two galliform species, Japanese Quail and Chicken, between sexes and between urine and faecal parts of droppings. Journal of Ornithology153(4), 1179–1188.

Loretto, M.-C., Fraser, O. N., & Bugnyar, T. (2012). Ontogeny of Social Relations and Coalition Formation in Common Ravens (Corvus corax). International Journal of Comparative Psychology25(3), 180–194.

Schloegl, C., Schmidt, J., Boeckle, M., Weiß, B. M., & Kotrschal, K. (2012). Grey parrots use inferential reasoning based on acoustic cues alone. Proceedings. Biological Sciences279(1745), 4135–4142.

Schwab, C., Swoboda, R., Kotrschal, K., & Bugnyar, T. (2012). Recipients affect prosocial and altruistic choices in jackdaws, Corvus monedula. PLoS ONE7(4), [e34922].

Wascher, C. A. F., Bauer, A. C., Holtmann, A. R., & Kotrschal, K. (2012). Environmental and social factors affecting the excretion of intestinal parasite eggs in graylag geese. Behavioral Ecology23(6), 1276–1283.

Wascher, C. A. F., Dufour, V., & Bugnyar, T. (2012). Carrion crows cannot overcome impulsive choice in a quantitative exchange task. Frontiers in Psychology3, [118].

Wascher, C. A. F., Szipl, G., Boeckle, M., & Wilkinson, A. (2012). You sound familiar: carrion crows can differentiate between the calls of known and unknown heterospecifics. Animal Cognition15(5), 1015–1019.

Wascher, C. A. F., Weiss, B. M., Arnold, W., & Kotrschal, K. (2012). Physiological implications of pair-bond status in greylag geese. Biology Letters8(3), 347–350.



Beetz, A., Kotrschal, K., Turner, D. C., Hediger, K., Uvnäs-Moberg, K., & Julius, H. (2011). The effect of a real dog, toy dog and friendly person on insecurely attached children during a stressful task: An exploratory study. Anthrozoös24(4), 349–368.

Bugnyar, T. (2011). Knower-guesser differentiation in ravens: others’ viewpoints matter. Proceedings. Biological Sciences278(1705), 634–640.

Fraser, O. N., & Bugnyar, T. (2011). Ravens reconcile after aggressive conflicts with valuable partners. PLoS ONE6(3), [e18118].

Kehmeier, S., Schloegl, C., Scheiber, I. B. R., & Weiss, B. M. (2011). Early development of gaze following into distant space in juvenile Greylag geese (Anser anser). Animal Cognition14(4), 477–485.

Kenward, B., Schloegl, C., Rutz, C., Weir, A. A. S., Bugnyar, T., & Kacelnik, A. (2011). On the evolutionary and ontogenetic origins of tool-oriented behaviour in New Caledonian crows (Corvus moneduloides). Biological Journal of the Linnean Society102(4), 870–877.

Mikolasch, S., Kotrschal, K., & Schloegl, C. (2011). African grey parrots (Psittacus erithacus) use inference by exclusion to find hidden food. Biology Letters7(6), 875–877.

Mikolasch, S., Kotrschal, K., & Schloegl, C. (2011). Is caching the key to exclusion in corvids? The case of carrion crows (Corvus corone corone). Animal Cognition15(1), 73–82.

Pika, S., & Bugnyar, T. (2011). The use of referential gestures in ravens (Corvus corax) in the wild. Nature Communications2, [560].

Scheiber, I. B. R., Hohnstein, A., Kotrschal, K., & Weiss, B. M. (2011). Juvenile greylag geese (Anser anser) discriminate between individual siblings. PLoS ONE6(8), [e22853].

Schloegl, C. (2011). What you see is what you get—Reloaded: Can jackdaws (Corvus monedula) find hidden food through exclusion? Journal of Comparative Psychology125(2), 162–174.

Schmidt, J., Scheid, C., Kotrschal, K., Bugnyar, T., & Schloegl, C. (2011). Gaze direction – A cue for hidden food in rooks (Corvus frugilegus)? Behavioural Processes88(2), 88–93.

Wagner, J., Szipl, G., & Schwab, C. (2011). Successfully releasing Jackdaws, Corvus monedula: spatial dispersion and the fusion of social groups. Die Vogelwarte49, 163–174.

Wedl, M., Bauer, B., Gracey, D., Grabmayer, C., Spielauer, E., Day, J., & Kotrschal, K. (2011). Factors influencing the temporal patterns of dyadic behaviours and interactions between domestic cats and their owners. Behavioural Processes86(1), 58–67.

Weiß, B. M., Kotrschal, K., & Foerster, K. (2011). A longitudinal study of dominance and aggression in greylag geese (Anser anser). Behavioral Ecology22(3), 616–624.



Fraser, O. N., & Bugnyar, T. (2010). Do ravens show consolation? Responses to distressed others. PLoS ONE5(5), [e10605].

Fraser, O. N., & Bugnyar, T. (2010). The quality of social relationships in ravens. Animal Behaviour79(4), 927–933.

Hemetsberger, J., Scheiber, I. B. R., Weiß, B. M., Frigerio, D., & Kotrschal, K. (2010). Influence of socially involved hand-raising on life history and stress responses in greylag geese. Interaction Studies11(3), 380–395.

Hirschenhauser, K., Weiß, B. M., Haberl, W., Möstl, E., & Kotrschal, K. (2010). Female androgen patterns and within-pair testosterone compatibility in domestic geese (Anser domesticus). General and Comparative Endocrinology165(2), 195–203.

Kotrschal, K., Scheiber, I. B. R., & Hirschenhauser, K. (2010). Individual performance in complex social systems: the greylag goose example BT - Animal Behaviour: Evolution and Mechanisms. In P. Kappeler (Ed.), Animal Behaviour: Evolution and Mechanisms (pp. 121–148). Springer Berlin Heidelberg.

Kralj-Fišer, S., Scheiber, I. B. R., Kotrschal, K., Weiß, B. M., & Wascher, C. A. F. (2010). Glucocorticoids enhance and suppress heart rate and behaviour in time dependent manner in greylag geese (Anser anser). Physiology & Behavior100(4), 394–400.

Kralj-Fišer, S., Weiß, B. M., & Kotrschal, K. (2010). Behavioural and physiological correlates of personality in greylag geese (Anser anser). Journal of Ethology28(2), 363–370.

Loretto, M.-C., Schloegl, C., & Bugnyar, T. (2010). Northern bald ibises follow others’ gaze into distant space but not behind barriers. Biology Letters6(1), 14-17.

Scheiber, I. B. R. (2010). Sozialverhalten im Lichte der Evolutionstheorie und Verhaltensbiologie. In R. Neck (Hrsg), Evolution: Natur - Mensch - Gesellschaft. Peter Lang GmbH.

Wascher, C. A. F., Fraser, O. N., & Kotrschal, K. (2010). Heart rate during conflicts predicts post-conflict stress-related behavior in Greylag geese. PLoS ONE5(12), [e15751].

Wedl, M., Schöberl, I., Bauer, B., Day, J., & Kotrschal, K. (2010). Relational factors affecting dog social attraction to human partners. Interaction Studies11(3), 482–503.

Weiß, B. M., Kehmeier, S., & Schloegl, C. (2010). Transitive inference in free-living greylag geese, Anser anser. Animal Behaviour79(6), 1277–1283.

Weiß, B. M., Kotrschal, K., Möstl, E., & Hirschenhauser, K. (2010). Social and life-history correlates of hormonal partner compatibility in greylag geese (Anser anser). Behavioral Ecology21(1), 138–143.



Bertin, A., Hirschenhauser, K., & Kotrschal, K. (2009). Trans-Generational Influence of Human Disturbances in Japanese Quail: Egg Quality Influences Male Social and Sexual Behaviour. Ethology115(9), 879–887.

Bertin, A., Richard-Yris, M. A., Houdelier, C., Richard, S., Lumineau, S., Kotrschal, K., & Möstl, E. (2009). Divergent selection for inherent fearfulness leads to divergent yolk steroid levels in quail. Behaviour146(6), 757–770.

Day, J.E.L., Kergoat, S., & Kotrschal, K. (2009). Do pets influence the quantity and choice of food offered to them by their owners: lessons from other animals and the pre-verbal human infant? CAB Reviews: Perspectives in Agriculture, Veterinary Science, Nutrition and Natural Resources4(42), 1–12.

Kotrschal, K. (2009). Die evolutionäre Theorie der Mensch-Tier-Beziehung. in C. Otterstedt, & M. Rosenberger (Hrsg.), Gefährten - Konkurrenten - Verwandte. Die Mensch-Tier-Beziehung im wissenschaftlichen Diskurs (S. 88-77). Vandenhoeck & Ruprecht.

Kotrschal, K., Schöberl, I., Bauer, B., Thibeaut, A.-M., & Wedl, M. (2009). Dyadic relationships and operational performance of male and female owners and their male dogs. Behavioural Processes81(3), 383–391.

Scheiber, I. B. R., Kotrschal, K., & Weiss, B. M. (2009a). Benefits of family reunions: Social support in secondary Greylag goose families. Hormones and Behavior55(1), 133–138.

Scheiber, I. B. R., Kotrschal, K., & Weiss, B. M. (2009b). Serial agonistic attacks by Greylag goose families, Anser anser, against the same opponent. Animal Behaviour77(5), 1211–1216.

Schloegl, C., Bugnyar, T., & Aust, U. (2009). Exclusion performances in non-human animals: from pigeons to chimpanzees and back again. In S. Watanabe, A. P. Blaisdell, L. Huber & A. Young (eds), Rational Animals, Irrational Humans (pp. 217-234). Keio University Press, Tokyo

Schloegl, C., Dierks, A., Gajdon, G. K., Huber, L., Kotrschal, K., & Bugnyar, T. (2009). What you see is what you get? Exclusion performances in ravens and keas. PLoS ONE4(8), [e6368].

Wascher, C. A. F., Scheiber, I. B. R., Weiß, B. M., & Kotrschal, K. (2009). Heart rate responses to agonistic encounters in greylag geese, Anser anser. Animal Behaviour77(4), 955–961.

Wedl, M., Bauer, B., Dittami, J., Schöberl, I., & Kotrschal, K. (2009). Effects of personality and sex on behavioral patterns in human--dog dyads. Journal of Veterinary Behavior: Clinical Applications and Research4(2), 88–89.

Wedl, M., & Kotrschal, K. (2009). Social and Individual Components of Animal Contact in Preschool Children. Anthrozoös22(4), 383–396.



Bertin, A., Richard-Yris, M.-A., Houdelier, C., Lumineau, S., Möstl, E., Kuchar, A., Hirschenhauser, K., & Kotrschal, K. (2008). Habituation to humans affects yolk steroid levels and offspring phenotype in quail. Hormones and Behavior54(3), 396–402.

Bókony, V., Garamszegi, L. Z., Hirschenhauser, K., & Liker, A. (2008). Testosterone and melanin-based black plumage coloration: a comparative study. Behavioral Ecology and Sociobiology62(8), 1129-1238.

Bugnyar, T. (2008). Animal Cognition: Rooks Team up to Solve a Problem. Current Biology18(12), R530–R532.

Garamszegi, L. Z., Hirschenhauser, K., Bókony, V., Eens, M., Hurtrez‐Boussès, S., Møller, A. P., Oliveira, R., & Wingfield, J. C. (2008). Latitudinal Distribution, Migration, and Testosterone Levels in Birds. The American Naturalist172(4), 533–546.

Hirschenhauser, K., Canário, A., Ros, A., Taborsky, M., & Oliveira, R. (2008). Social context may affect urinary excretion of 11-ketotestosterone in African cichlids. Behaviour145(10), 1367–1388.

Hirschenhauser, K., Wittek, M., Johnston, P., & Möstl, E. (2008). Social context rather than behavioral output or winning modulates post-conflict testosterone responses in Japanese quail (Coturnix japonica). Physiology & Behavior95(3), 457–463.

Kotrschal, K. (2008). Angepasst woran? Vom Mängel- zum Konfliktwesen. in H. Schmidinger, & C. Sedmak (Hrsg.), Der Mensch - Ein Mängelwesen? Endlichkeit-Kompensation-Entwicklungen (S. 55-65). Wissenschaftliche Buchgesellschaft.

Kotrschal, K. (2008). Geist als Anpassung? in K. Appel, H. Philipp, R. Langthaler, & S. Müller (Hrsg.), Naturalisierung des Geistes?: Beiträge zur Gegenwärtigen Debatte um den Geist (S. 84-92). Königshausen und Neumann.

Range, F., Bugnyar, T., & Kotrschal, K. (2008). The performance of ravens on simple discrimination tasks: a preliminary study. Acta Ethologica11(1), 34–41.

Scheiber, I. B. R., Weiss, B.M., Hirschenhauser, K., Wascher, C. A. F., Nedelcu, I. T., & Kotrschal, K. (2008). Does “relationship intelligence” make big brains in birds? The Open Biology Journal1, 6–8.

Scheid, C., & Bugnyar, T. (2008). Short-term observational spatial memory in Jackdaws (Corvus monedula) and Ravens (Corvus corax). Animal Cognition11(4), 691–698.

Scheid, C., Schmidt, J., & Noë, R. (2008). Distinct patterns of food offering and co-feeding in rooks. Animal Behaviour76(5), 1701–1707.

Schloegl, C., Kotrschal, K., & Bugnyar, T. (2008a). Do common ravens (Corvus corax) rely on human or conspecific gaze cues to detect hidden food? Animal Cognition11(2), 231–241.

Schloegl, C., Kotrschal, K., & Bugnyar, T. (2008b). Modifying the object-choice task: Is the way you look important for ravens? Behavioural Processes77(1), 61–65.

Schlögl, H. C., Schmidt, J., Scheid, C., Kotrschal, K., & Bugnyar, T. (2008). Gaze following in non-human animals: The corvid example. in E. A. Weber, & L. H. Krause (Eds.), Animal Behaviour: New Research (S. 73-92). Nova Science Publishers.

Scholl, S., Grall, G., Petzl, V., Röthler, M., Slotta-Bachmayr, L., & Kotrschal, K. (2008). Behavioural effects of goats on disabled persons. Therapeutic Communities29(3), 297–309.

Schwab, C., Bugnyar, T., & Kotrschal, K. (2008). Preferential learning from non-affiliated individuals in jackdaws (Corvus monedula). Behavioural Processes79(3), 148–155.

Schwab, C., Bugnyar, T., Schloegl, C., & Kotrschal, K. (2008). Enhanced social learning between siblings in common ravens, Corvus corax. Animal Behaviour75(2), 501–508.

Scott, A., Hirschenhauser, K., Bender, N., Oliveira, R., Earley, R., Sebire, M., Ellis, T., Pavlidis, M., Hubbard, P.C., Huertas, M., & Canario, A. (2008). Non-invasive measurement of steroids in fish-holding water: important considerations when applying the procedure to behaviour studies. Behaviour145(10), 1307–1328.

Stöwe, M., Bugnyar, T., Schloegl, C., Heinrich, B., Kotrschal, K., & Möstl, E. (2008). Corticosterone excretion patterns and affiliative behavior over development in ravens (Corvus corax). Hormones and Behavior53(1), 208–216.

Wascher, C. A. F., Arnold, W., & Kotrschal, K. (2008). Heart rate modulation by social contexts in Greylag geese (Anser anser). Journal of Comparative Psychology122(1), 100–107.

Wascher, C. A. F., Scheiber, I. B. R., & Kotrschal, K. (2008). Heart rate modulation in bystanding geese watching social and non-social events. Proceedings. Biological Sciences275(1643), 1653–1659.

Weiß, B. M., Kotrschal, K., Frigerio, D., Hemetsberger, J., & Scheiber, I. (2008). Birds of a feather stay together: Extended family bonds, clan structures and social support in greylag geese (Anser anser). in R. N. Ramirez (Hrsg.), Family Relations: Issues and Challenges (S. 69-88). Nova Science Publishers.

Weiß, B. M., Poggemann, K., Olek, K., Foerster, K., & Hirschenhauser, K. (2008). Isolation and characterization of microsatellite marker loci in the greylag goose (Anser anser). Molecular Ecology Resources8(6), 1411–1413.



Bugnyar, T. (2007). An integrative approach to the study of "theory-of-mind"-like abilities in ravens. Japanese Journal of Animal Psychology, 57(1), 15–27.

Bugnyar, T., Schwab, C., Schloegl, C., Kotrschal, K., & Heinrich, B. (2007). Ravens Judge Competitors through Experience with Play Caching. Current Biology17(20), 1804–1808.

Bugnyar, T., Stöwe, M., & Heinrich, B. (2007). The ontogeny of caching in ravens, Corvus corax. Animal Behaviour74(4), 757–767.

Schloegl, C., Kotrschal, K., & Bugnyar, T. (2007). Lektionen von Rabenvögeln und Gänsen. Instinkt oder Verstand? Biologie in Unserer Zeit37(6), 366–374.

Kralj-Fišer, S., Scheiber, I. B. R., Blejec, A., Moestl, E., & Kotrschal, K. (2007). Individualities in a flock of free-roaming greylag geese: Behavioral and physiological consistency over time and across situations. Hormones and Behavior51(2), 239–248.

Scheid, C., Range, F., & Bugnyar, T. (2007). When, What, and Whom to Watch? Quantifying Attention in Ravens (Corvus corax) and Jackdaws (Corvus monedula). Journal of Comparative Psychology121(4), 380–386.

Schloegl, C., Kotrschal, K., & Bugnyar, T. (2007). Gaze following in common ravens, Corvus corax: ontogeny and habituation. Animal Behaviour74(4), 769–778.

Stöwe, M., & Kotrschal, K. (2007). Behavioural phenotypes may determine whether social context facilitates or delays novel object exploration in ravens (Corvus corax). Journal of Ornithology148(2), 179–184.

Zoufal, K., Fritz, J., Bichler, M., Kirbauer, M., Markut, T., Meran, I., Wolf, A., & Kotrschal, K. (2007). Feeding ecology of the Northern Bald Ibis in different habitat types: an experimental field study with hand-raised individuals. In C. Bowden, & C. Böhm (Eds.), Report of the 2nd IAGNBI Meeting 2nd IAGNBI Meeting (pp. 77-84). Vejer.



Bugnyar, T., & Heinrich, B. (2006). Pilfering ravens, Corvus corax, adjust their behaviour to social context and identity of competitors. Animal Cognition9(4), 369–376.

Hirschenhauser, K., & Oliveira, R. F. (2006). Social modulation of androgens in male vertebrates: meta-analyses of the challenge hypothesis. Animal Behaviour71(2), 265–277.

Kotrschal, K., Hemetsberger, J., & Weiß, B. M. (2006). Making the best of a bad situation: Homosociality in male greylag geese. In V. Sommer (Ed.), Homosexual behaviour in animals: An evolutionary perspective (pp. 45-76). Cambridge Univ. Press.

Range, F., Bugnyar, T., Schlögl, C., & Kotrschal, K. (2006). Individual and sex differences in learning abilities of ravens. Behavioural Processes73(1), 100–106.

Sorato, E., & Kotrschal, K. (2006). Hormonal and behavioural symmetries between the sexes in the Northern bald ibis. General and Comparative Endocrinology146(3), 265–274.

Stöwe, M., Bugnyar, T., Heinrich, B., & Kotrschal, K. (2006). Effects of Group Size on Approach to Novel Objects in Ravens (Corvus corax). Ethology112(11), 1079–1088.

Stöwe, M., Bugnyar, T., Loretto, M.-C., Schloegl, C., Range, F., & Kotrschal, K. (2006). Novel object exploration in ravens (Corvus corax): Effects of social relationships. Behavioural Processes73(1), 68–75.



Bugnyar, T., Stöwe, M., & Heinrich, B. (2005). Ravens, Corvus corax, follow gaze direction of humans around obstacles. Proceedings. Biological Sciences271(1546).

Heinrich, B., & Bugnyar, T. (2005). Testing Problem Solving in Ravens: String-Pulling to Reach Food. Ethology111(10), 962–976.

Hemetsberger, J. (2005). Frühe Brutnachweise bei Steinlerche Ammomanes isabellina, Saharasteinschmätzer Oenanthe leucopyga, Akaziendrossling Turdoides fulvus und Hausammer Emberiza striolata in der zentralen Sahara Algeriens. Vogelwarte, 43, 271.

Hirschenhauser, K., & Frigerio, D. (2005). 5. Hidden Patterns of Male Sex Hormones and Behaviour Vary with Life History. In G. R. L. Anolli, S. Duncan, M. S. Magnusson (Ed.), The hidden structure of social interaction. Amsterdam: Citeseer.

Hirschenhauser, K., Kotrschal, K., & Möstl, E. (2005). Synthesis of Measuring Steroid Metabolites in Goose Feces. Annals of the New York Academy of Sciences1046(1), 138–153.

Niall Daisley, J., Bromundt, V., Möstl, E., & Kotrschal, K. (2005). Enhanced yolk testosterone influences behavioral phenotype independent of sex in Japanese quail chicks Coturnix japonica. Hormones and Behavior47(2), 185–194.

Scheiber, I. B. R., Kralj, S., & Kotrschal, K. (2005). Sampling Effort/Frequency Necessary to Infer Individual Acute Stress Responses from Fecal Analysis in Greylag Geese (Anser anser). Annals of the New York Academy of Sciences1046(1), 154–167.

Scheiber, I. B. R., Weiss, B. M., Frigerio, D., & Kotrschal, K. (2005). Active and passive social support in families of Greylag geese (Anser anser). Behaviour142, 1535–1557.



Bugnyar, T., & Kotrschal, K. (2004). Leading a conspecific away from food in ravens (Corvus corax)? Animal Cognition7(2), 69–76.

Bugnyar, T., Stöwe, M., & Heinrich, B. (2004). Ravens, Corvus corax, follow gaze direction of humans around obstacles. Proceedings of the Royal Society of London. Series B: Biological Sciences271(1546).

Frigerio, D., Dittami, J., Möstl, E., & Kotrschal, K. (2004). Excreted corticosterone metabolites co-vary with ambient temperature and air pressure in male Greylag geese (Anser anser). General and Comparative Endocrinology137(1), 29–36.

Frigerio, D., Hirschenhauser, K., Möstl, E., Dittami, J., & Kotrschal, K. (2004). Experimentally elevated testosterone increases status signalling in male Greylag geese (Anser anser). Acta Ethologica7(1), 9–18.

Goymann, W., Moore, I. T., Scheuerlein, A., Hirschenhauser, K., Grafen, A., & Wingfield, J. C. (2004). Testosterone in Tropical Birds: Effects of Environmental and Social Factors. The American Naturalist164(3), 327–334.

Hemetsberger, J., & Petutschnig, B. (2004). Zur Avifauna Jordaniens. Denisia, 14, 421-436.

Hirschenhauser, K., Taborsky, M., Oliveira, T., Canàrio, A. V. M., & Oliveira, R. F. (2004). A test of the “challenge hypothesis” in cichlid fish: simulated partner and territory intruder experiments. Animal Behaviour68(4), 741–750.

Weiß, B. M., & Kotrschal, K. (2004). Effects of Passive Social Support in Juvenile Greylag Geese (Anser anser): A Study from Fledging to Adulthood. Ethology110(6), 429–444.



Frigerio, D., Weiss, B., Dittami, J., & Kotrschal, K. (2003). Social allies modulate corticosterone excretion and increase success in agonistic interactions in juvenile hand-raised graylag geese (Anser anser). Canadian Journal of Zoology81(10), 1746–1754.

Hackl, R., Bromundt, V., Daisley, J., Kotrschal, K., & Möstl, E. (2003). Distribution and origin of steroid hormones in the yolk of Japanese quail eggs (Coturnix coturnix japonica). Journal of Comparative Physiology B173(4), 327–331.

Hirschenhauser, K., Winkler, H., & Oliveira, R. F. (2003). Comparative analysis of male androgen responsiveness to social environment in birds: the effects of mating system and paternal incubation. Hormones and Behavior43(4), 508–519.

Kotrschal, K. (2003). Das Ende der Beliebigkeit — zum evolutionär-biologischen Fundament des Menschen. Österreichische Zeitschrift Für Soziologie28(2), 21–43.

Kotrschal, K., & Ortbauer, B. (2003). Behavioral effects of the presence of a dog in a classroom. Anthrozoös16(2), 147–159.



Bugnyar, T., & Kotrschal, K. (2002). Observational learning and the raiding of food caches in ravens, Corvus corax: is it “tactical” deception? Animal Behaviour64(2), 185–195.

Bugnyar, T., & Kotrschal, K. (2002). Scrounging Tactics in Free-Ranging Ravens, Corvus corax. Ethology108(11), 993–1009.

Fritz, J., & Kotrschal, K. (2002). On avian imitation: cognitive and ethological perspectives. In Nehaniv, C. L., Dautenhahn, K., & Dautenhan, K. Imitation in Animals and Artifact (p. 133–155). Bradford Books

Hirschenhauser, K., Frigerio, D., Grammer, K., & Magnusson, M. S. (2002). Monthly Patterns of Testosterone and Behavior in Prospective Fathers. Hormones and Behavior42(2), 172–181.

Kijne, M., & Kotrschal, K. (2002). Neophobia affects choice of food-item size in group-foraging common ravens (Corvus corax). Acta Ethologica5(1), 13–18.

Kotrschal, K. (2002). Eine integrative, systematische Sicht sozialer. In U. Ganßloser (Ed.), Soziale Mechanismen. Filander Verlag.

Pfeffer, K., Fritz, J., & Kotrschal, K. (2002). Hormonal correlates of being an innovative greylag goose, Anser anser. Animal Behaviour63(4), 687–695.

Tintner, A., & Kotrschal, K. (2002). Early social influence on nestling development in Waldrapp ibis (Geronticus eremita). Zoo Biology21(5), 467–480.



Bugnyar, T., Kijne, M., & Kotrschal, K. (2001). Food calling in ravens: are yells referential signals? Animal Behaviour61(5), 949–958.

Bugnyar, T., & Kotrschal, K. (2001). Movement coordination and signalling in ravens (Corvus corax): an experimental field study. Acta Ethologica3(2), 101–109.

Frigerio, D., Moestl, E., & Kotrschal, K. (2001). Excreted Metabolites of Gonadal Steroid Hormones and Corticosterone in Greylag Geese (Anser anser) from Hatching to Fledging. General and Comparative Endocrinology124(2), 246–255.

Frigerio, D., Weiss, B., & Kotrschal, K. (2001). Spatial proximity among adult siblings in Greylag geese (Anser anser): evidence for female bonding? Acta Ethologica3(2), 121–125.

Fritz, J. (2001). Der intelligente Kumpan am Beispiel der Graugänse. In K. Kotrschal, G. Müller, & H. Winkler (Eds.), Konrad Lorenz und seine verhaltensbiologischen Konzepte aus heutiger Sicht (p. 261-270). Filander Verlag.

Hemetsberger, J. (2001). Die Entwicklung der Grünauer Graugansschar seit 1973. In K. Kotrschal, G. Müller, & H. Winkler (Eds.), Konrad Lorenz und seine verhaltensbiologischen Konzepte aus heutiger Sicht (p. 249-260). Filander Verlag.

Kotrschal, K. (2001). Das Rousseausche Vorurteil vom „edlen Wilden “: Die Wurzeln der Ethik von Konrad Lorenz. In K. Kotrschal, G. Müller, & H. Winkler (Eds.), Konrad Lorenz und seine verhaltensbiologischen Konzepte aus heutiger Sicht (p. 109-118). Filander Verlag.

Kotrschal, K. (2001). Der Beitrag von Österreich zur Wissenschaft der Biologie vom Verhalten und zum evolutionären Bild des Menschen. In K. Acham (Ed.), Cognitio Humana. Zur Geistesgeschichte Österreichs.

Kotrschal, K., Bugnyar, T., & Stöwe, M. (2001). Kognition und Neophobie bei Raben. Charandrius3, 127–135.

Mostl, E., Spendier, H., & Kotrschal, K. (2001). Concentration of immunoreactive progesterone and androgens in the yolk of hens’ eggs (Gallus domesticus). Wiener Tierarztliche Monatsschrift88(3), 62–65.


Older Publications

Fritz, J., Bisenberger, A., & Kotrschal, K. (2000). Stimulus enhancement in greylag geese: socially mediated learning of an operant task. Animal Behaviour59(6), 1119–1125.

Hemetsberger, J. (2000). Erste Ergebnisse des Jahres 1999. Vogelkundliche Nachrichten Oberösterreich - Naturschutz Aktuell8(1), 19–27.

Hirschenhauser, K., Möstl, E., Péczely, P., Wallner, B., Dittami, J., & Kotrschal, K. (2000). Seasonal Relationships between Plasma and Fecal Testosterone in Response to GnRH in Domestic Ganders. General and Comparative Endocrinology118(2), 262–272.

Hirschenhauser, K., Möstl, E., Wallner, B., Dittami, J., & Kotrschal, K. (2000). Endocrine and Behavioural Responses of Male Greylag Geese (Anser anser) to Pairbond Challenges during the Reproductive Season. Ethology106(1), 63–77.

Kotrschal, K. (2000). Taste(s) and olfaction(s) in fish: a review of spezialized sub-systems and central integration. Pflügers Archiv - European Journal of Physiology439(7), R178–R180.

Kotrschal, K., Dittami, J., Hirschenhauser, K., Möstl, E., & Peczely, P. (2000). Effects of physiological and social challenges in different seasons on fecal testosterone and corticosterone in male domestic geese (Anser domesticus). Acta Ethologica2(2), 115–122.

Fritz, J., & Kotrschal, K. (1999). Social learning in common ravens, Corvus corax. Animal Behaviour57(4), 785–793.

Hirschenhauser, K., Mostl, E., & Kotrschal, K. (1999). Within-pair testosterone covariation and reproductive output in Greylag Geese Anser anserIbis141(4), 577–586.

Hirschenhauser, K., Möstl, E., & Kotrschal, K. (1999). Seasonal Patterns of Sex Steroids Determined from Feces in Different Social Categories of Greylag Geese (Anser anser). General and Comparative Endocrinology114(1), 67–79.

Kotrschal, K. (1998). Sensory Systems. In M. Horn, M. Martin, & K. Chotkowski (Ed.), Intertidal Fishes (pp. 126–142). Academic Press.

Kotrschal, K., Hirschenhauser, K., & Möstl, E. (1998). The relationship between social stress and dominance is seasonal in greylag geese. Animal Behaviour55(1), 171–176.

Kotrschal, K., Van Staaden, M. J., & Huber, R. (1998). Fish Brains: Evolution and environmental Relationships. Reviews in Fish Biology and Fisheries8(4), 373–408.

Kotrschal, K. (1997). Die Konrad Lorenz Forschungsstelle in Grünau. Vogelkundliche Nachrichten Oberösterreich - Naturschutz Aktuell4, 31–33.

Kotrschal, K., Krautgartner, W.-D., & Hansen, A. (1997). Ontogeny of the Solitary Chemosensory Cells in the Zebrafish, Danio rerioChemical Senses22(2), 111–118.

Kotrschal, K. (1996). Solitary chemosensory cells: why do primary aquatic vertebrates need another taste system? Trends in Ecology & Evolution11(3), 110–114.

Kotrschal, K., Peters, R. C., & Døving, K. B. (1996). Chemosensory and tactile nerve responses from the anterior dorsal fin of a rockling, Gaidropsarus vulgaris (Gadidae, Teleostei). Primary Sensory Neuron1, 297–310.

Drack, G., & Kotrschal, K. (1995). Aktivitätsmuster und Spiel von freilebenden Kolkraben Corvus corax im inneren Almtal/Oberösterreich. Monticula7, 159–174.

Kotrschal, K. (1995). Ecomorphology of solitary chemosensory cell systems in fish: a review. Environmental Biology of Fishes44(1), 143–155.

Kotrschal, K., & Essler, H. (1995). Goals and approaches in the analysis of locomotion in fish, with a focus on laboratory studies. Reviews in Fisheries Science3(2), 171–200.

Essler, H., & Kotrschal, K. (1994). High resolution analysis of swim path patterns of intact and olfaction-deprived minnows (Phoxinus phoxinus) stimulated with food and potential predator odour. Journal of Fish Biology45(4), 555–567.

Kotrschal, K., Peters, R., & Atema, J. (1994). Zoology: analysis of complex systems: ZACS. Zoologische Jahrbücher. Abteilung für allgemeine Zoologie und Physiologie der Tiere, 97(1). G. Fischer.

Kotrschal, K., Hemetsberger, J., & Dittami, J. (1993). Food exploitation by a winter flock of greylag geese: behavioral dynamics, competition and social status. Behavioral Ecology and Sociobiology33(5), 289–295.

Kotrschal, K., Peters, R., & Atema, J. (1993). Sampling and behavioral evidence for mucus detection in a unique chemosensory organ: the anterior dorsal fin in rocklings (Ciliata mustela: Gadidae: Teleostei). Zoologische Jahrbücher. Abteilung Für Allgemeine Zoologie Und Physiologie Der Tiere97(1), 47–67.

Kotrschal, K., Whitear, M., & Finger, T. E. (1993). Spinal and facial innervation of the skin in the gadid fish Ciliata mustela (Teleostei). The Journal of Comparative Neurology331(3), 407–417.

Kotrschal, K. (1992). Quantitative scanning electron microscopy of solitary chemoreceptor cells in cyprinids and other teleosts. Environmental Biology of Fishes35(3), 273–282.

Kotrschal, K., Hemetsberger, J., & Dittami, J. (1992). Vigilance in a flock of semi-tame Greylag Geese Answer answer in response to approaching eagles Haliaeetus albicilla and Aquila chrysaetos. Wildfowl43(43), 215–219.

Kotrschal, K., & Palzenberger, M. (1992). Neuroecology of cyprinids: comparative, quantitative histology reveals diverse brain patterns. In W. Wieser, F. Schiemer, A. Goldschmidt, K. Kotrschal (eds) Environmental biology of European cyprinids. Developments in environmental biology of fishes, vol 13. Springer, Dordrecht.

Mausz, B., Dittami, J., Kotrschal, K. (1992). Triumphgeschrei und Aggression bei der Graugans (Anser anser). Ökologie Der Vögel14, 165–172.

Finger, T. E., Drake, S. K., Kotrschal, K., Womble, M., & Dockstader, K. C. (1991). Postlarval growth of the peripheral gustatory system in the channel catfish, Ictalurus punctatus. The Journal of Comparative Neurology314(1), 55–66.

Kotrschal, K. (1991). Solitary chemosensory cells — taste, common chemical sense or what? Reviews in Fish Biology and Fisheries1(1), 3–22.

Peters, R. C., Kotrschal, K., & Krautgartner, W.-D. (1991). Solitary chemoreceptor cells of Ciliata mustela (Gadidae, Teleostei) are tuned to mucoid stimuli. Chemical Senses16(1), 31–42.

Kotrschal, K. (1989). Trophic ecomorphology in eastern Pacific blennioid fishes: character transformation of oral jaws and associated change of their biological roles. Environmental Biology of Fishes24(3), 199–218.

Kotrschal, B. K. (1988). Evolutionary patterns in tropical marine reef fish feeding. Journal of Zoological Systematics and Evolutionary Research26(1), 51–64.

Kotrschal, K., & Whitear, M. (1988). Chemosensory anterior dorsal fin in rocklings (Gaidropsarus and Ciliata, Teleostei, Gadidae): Somatotopic representation of the ramus recurrens facialis a revealed by transganglionic transport of HRP. The Journal of Comparative Neurology268(1), 109–120.

Lindquist, D. G., & Kotrschal, K. M. (1987). The Diets in Four Pacific Tube Blennies (Acanthemblemaria: Chaenopsidae): Lack of Ecological Divergence in Syntopic Species. Marine Ecology8(4), 327–335.

Kotrschal, K., & Lindquist, D. G. (1986). The Feeding Apparatus in four Pacific Tube Blennies (Teleostei: Chaenopsidae): Marine Ecology7(3), 241–253.

Goldschmid, A., & Kotrschal, K. (1985). Morphological and functional adaptations in different feeding types of blennies (Blenniidae, Perciformes, Teleostei). Fortschritte der Zoologie30, 241–244.