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Here you can find a list of the publications of the KLF (in chronological order).



  • Frigerio, D., Cibulski, L., Ludwig, S. C., Campderrich, I., Kotrschal, K. & Wascher, C. A. F. (2016): Excretion patterns of coccidian oocysts and nematode eggs during the reproductive season in Northern Bald Ibis (Geronticus eremita). Journal of Ornithology, in press.


  • Loretto, M.-C., R. Schuster and T. Bugnyar. (2015): GPS tracking of non-breeding ravens reveals importance of anthropogenic food sources during their dispersal in the Eastern Alps. Current Zoology in press. 
  • Loretto, M.-C., R. Schuster, S. Reimann, D.M. Graulich and T. Bugnyar. (2015): Shared space - individually used: Spatial behaviour of non-breeding ravens (Corvus corax) close to a permanent anthropogenic food source. Journal of Ornithology in press.
  • Szipl, G., Böckle, M., Wascher, C., Spreafico, M., & Bugnyar, T. (2015): With whom to dine? Ravens' responses to food-associated calls depend on individual characteristics of the caller. Animal Behaviour, 99(1), 33-42.


  • Dorn, S., Wascher, C. A. F., Möstl, E. & Kotrschal, K. (2014): Ambient temperature and air pressure modulate hormones and behaviour in Greylag geese (Anser anser) and Northern bald ibis(Geronticus eremita). Behavioural Processes 108: 27-35
  • Cibulski, L., Wascher, C., Weiß, B. M. & Kotrschal, K. (2014): Familiarity with the experimenter influences the performance of Common ravens (Corvus corax) and Carrion crows (Corvus corone corone) in cognitive tasks. Behavioural Processes. 103C, S. 129-137
  • Hillemann, F., Bugnyar, T., Kotrschal, K. & Wascher, C.A.F. (2014): Waiting for better, not for more: corvids respond to quality in two delay maintenance tasks. Animal Behaviour, 90, 1-10.
  • Massen, J.J.M., Pasukonis, A., Schmidt, J. & Bugnyar, T. (2014): Ravens notice dominance reversals among conspecifics within and outside their social group. Nature Communications 5, 3679.
  • Massen, J.J.M., Szipl, G. Spreafico, M. & Bugnyar, T. (2014): Ravens intervene in others’ bonding attempts. Current Biol., 24, 1-4.


  • Wascher, C.A.F., Bugnyar, T. (2013): Behavioral Responses to Inequity in Reward Distribution and Working Effort in Crows and Ravens; PLoS ONE 8(2): e56885. doi:10.1371/journal.pone.0056885, (journal link)
  • Guibert, F., Lumineau, S., Kotrschal, K., Möstl, E., Richard-Yris, M-A., Houdelier, C. (2013): Trans-generational effects of prenatal stress in quail. Proceedings of the Royal Society of London: Series B, Biological sciences 280 (1753): 1471-2954. 10.1098/rspb.2012.2368.
  • Frigerio,  D., Gegendorfer, G. (2013): Exotisch  oder heimisch? – Der Waldrapp  aus dem Almtal. Groß und Klein erforschen  die vom Aussterben  bedrohte  Vogelart.  Öko-L 35(3): 3-13.
  • Hirschenhauser, K., Gahr, M., Goymann, W. (2013): Winning and losing in public: Audiences direct future success in Japanese quail. Hormones and Behavior 63: 625-633. 10.1016/j.yhbeh.2013.02.010.
  • Kotrschal, K. (2013): The Quest for Understanding Social Complexity. Nova Acta Leopoldina  NF 111, 77–88.
  • Mikolasch, S., Kotrschal, K., Schloegl, C. (2013): Transitive inference in jackdaws (Corvus monedula). Behavioural Processes 92: 113-117. 10.1016/j.beproc.2012.10.017.
  • Teschke, I., Wascher, C.A.F., Scriba, M.F., von Bayern, A.M.P., Huml, V., Siemers, B., Tebbich, S. (2013) Did tool-use evolve with enhanced physical  cognitive abilities? Philosophical  Transactions of the Royal Society B 368: 20120418.
  • Ujfalussy, D. J., Miklósi, Á., Bugnyar, T., Kotrschal, K. (2013): Role of Mental Representations in Quantity Judgments by Jackdaws (Corvus monedula). Journal of Comparative Psychology. Advance online publication. doi: 10.1037/a0034063.


  • Bisenberger, A., Frigerio, D. (2012): Biene Maja zeigt uns ihre Welt - von einer Projektidee zur angewandten Methodik tiergestützter Vermittlungsarbeit mit Kleintieren; Öko-L (2): 29-35. 
  • Boeckle, M., Szipl G., Bugnyar, T. (2012): Who wants food? Individual characteristics in raven yells. Animal Behaviour, 84(5): 1123–1130 (journal link)
  • Boeckle, M., Bugnyar, T. (2012): Long-Term Memory for Affiliates in Ravens, Current Biology, 22(9): 801–806; doi:10.1016/j.cub.2012.03.023, (journal link)
  • Braun, A., Bugnyar, T. (2012): Social bonds and rank acquisition in raven nonbreeder aggregations, Animal Behaviour 84, 1507-1515.
  • Braun, A., Walsdorff, T., Fraser, O. N., Bugnyar, T. (2012): Socialized sub-groups in a temporary stable Raven flock? J. Ornithol. DOI 10.1007/s10336-011-0810-2.
  • Fraser, O. N., Bugnyar, T. (2012): Reciprocity of agonistic support in ravens. Animal Behaviour 83, 171-177, DOI: 10.1016/j.anbehav.2011.10.023 Initiates file download 
  • Frigerio, D., Kotrschal, K., Millesi, E. und Hemetsberger, J. 2012: Pupils observing and measuring Greylag goose behaviour: methods and results. invited contribution; International Journal for Cross-Disciplinary Subjects in Education -IJCDSE, 3 (4), 871-876. Initiates file download 
  • Hirschenhauser, K., Spreitzer, K., Lepschy, M., Kotrschal, K., Möstl, E. (2012): Excreted corticosterone metabolites differ between two galliform species, Japanese Quail and Chicken, between sexes and between urine and faecal parts of droppings. Journal of Ornithology. DOI 10.1007/s10336-012-0848-9
  • Hirschenhauser, K. (2012): Testosterone and partner compatibility: Evidence and emerging questions. Ethology, 118, 799-811. doi: 10.1111/j.1439-0310.2012.02087.x. Initiates file download 
  • Loretto, M.-C., Fraser, O. N., Bugnyar, T. (2012): Ontogeny of Social Relations and Coalition Formation in Common Ravens (Corvus corax). International Journal of Comparative Psychology, 25 (3), 180-194. Initiates file download 
  • Mikolasch, S., Kotrschal, K., Schloegl, C. (2012): The influence of local enhancement on choice performances in African grey parrots (Psittacus erithacus) and jackdaws (Corvus monedula); Journal of Comparative Psychology; in press
  • Mikolasch, S., Kotrschal, K., Schloegl, C. (2012): Is caching the key to exclusion in corvids? The case of carrion crows (Corvus corone corone); Animal Cognition, 15 (1), 73-82. doi 10.1007/s10071-011-0434-1. Initiates file download 
  • Schloegl, C., Schmidt, J., Boeckle, M., Weiß, B. M., Kotrschal, K. (2012): Grey parrots use inferential reasoning based on acoustic cues alone. Proceedings of the Royal Society of London B, 279, 4135-4142.   doi:10.1098/rspb.2012.1292 Initiates file download
  • Schwab, C., Swoboda, R., Kotrschal, K., Bugnyar, T. (2012): Recipients Affect Prosocial and Altruistic Choices in Jackdaws, Corvus monedula. PLoS ONE 7(4): e34922. doi:10.1371/journal.pone.0034922 (journal link)
  • Wascher, C.A.F., Weiß, B.M., Arnold, W., Kotrschal, K. (2012): Physiological implications of pair-bond status in greylag geese. Biol. Letters 8(3): 347–350.
  • Wascher, C.A.F., Dufour, V., Bugnyar, T. (2012): Carrion crows cannot overcome impulsive choice in a quantitative exchange task. Frontiers in Comparative Psychology 3:118. doi: 10.3389/fpsyg.2012.00118 (journal link)
  • Wascher C.A.F., Szipl G., Boeckle M., Wilkinson A. (2012): You sound familiar- Carrion crows can differentiate between the calls of known and unknown heterospecifics. Animal Cognition: doi 10.1007/s10071-012-0508-8
  • Wascher, C.A.F., Bauer, A.C., Holtmann, A.R., Kotrschal, K. (2012): Environmental and social factors affecting the excretion of intestinal parasite eggs in graylag geese. Behavioral Ecology. doi: 10.1093/beheco/ars113
  • Weiß, B.M., Scheiber, I.R.B. (2012): Long-term memory of hierarchical relationships in free-living greylag geese. Animal Cognition. published online: DOI 10.1007/s10071-012-0554-2


  • Beetz, A., Kotrschal, K., Turner, D., Hediger, K., Uvnäs-Moberg, K., Julius, H. (2011): The effect of social support by a dog in insecurely attached children: An exploratory study. Anthrozoös, 24 (4), 349-368
  • Bugnyar, T. (2010): Knower-guesser differentiation in ravens: others' viewpoints matter. Proceedings of the Royal Society of London B; 278, 634-640 doi:10.1098/rspb.2010.1514 
  • Dufour, V., Wascher, C.A.F., Braun, A., Miller, R., Bugnyar, T. (2011): Corvids can decide if a future exchange is worth waiting for. Biol. Lett. doi:10.1098/rsbl.2011.0726.
  • Fraser, O. N., Bugnyar, T. (2011): Ravens reconcile after aggressive conflicts with valuable partners. PLoS ONE 6(3) e18118, DOI: 10.1371/journal.pone.0018118 (journal link)
  • Kehmeier, S., Schloegl, C., Scheiber, I.R.B., Weiß, B.M.(2011): " Early development of gaze following into distant space in juvenile Greylag geese (Anser anser)“, Animal Cognition, 14 (4), 477-485
  • Kenward, B., Schloegl, C., Rutz, C., Weir, A.A.S., Bugnyar, T., Kacelnik, K. (2011): On the evolutionary and ontogenetic origins of tool-oriented behaviour in New Caledonian crows (Corvus moneduloides); Biological Journal of the Linnean Society 102: 870-877  Initiates file download
  • Mikolasch, S., Kotrschal, K., Schloegl, C.(2011): African grey parrots (Psittacus erithacus) use inference by exclusion to find hidden food; Biology letters; doi: 10.1098/rsbl.2011.0500.
  • Scheiber, I.R.B., Hohnstein, A., Kotrschal, K., Weiß, B.M. (2011): Juvenile Greylag Geese (Anser anser) discriminate between individual siblings. PLoS ONE 6(8): e22853. doi:10.1371/journal.pone.0022853 (journal link).
  • Schloegl, C. (2011): What You See Is What You Get—Reloaded: Can Jackdaws (Corvus monedula) Find Hidden Food Through Exclusion? Journal of Comparative Psychology 125/2, 162-174. Initiates file download 
  • Schmidt, J., Scheid, C., Kotrschal, K., Bugnyar, T., Schloegl, C.(2011): Gaze direction - A cue for hidden food in rooks (Corvus frugilegus)?, Behavioural Processes  88(2), 88-93.
  • Wascher, C.A.F., Scheiber, I.B.R., Braun, A., Kotrschal, K. (2011): Heart Rate Responses to Induced Challenge Situations in Greylag Geese (Anser anser). J. Comp. Psychol. 125/1, 116-119.
  • Wascher, C.A.F., Weiß, B.M., Arnold W., Kotrschal, K. (2011): Physiological implications of pair-bond status in greylag geese. Biology Letters, doi: 10.1098/rsbl.2011.0917.
  • Wedl, M., Bauer, B., Grabmayer, C., Gracey, D., Spielauer, E., Day, J., Kotrschal, K. (2011): Factors influencing the temporal patterns of dyadic behaviours and interactions between domestic cats and their owners. Behavioural Processes 86: 58-67
  • Weiß, B. M., Kotrschal, K., Foerster, K. (2011): A longitudinal study of dominance and aggression in greylag geese (Anser anser). Behav. Ecol., 22(3): 609-615. Initiates file download 


  • Fraser, O. N., Bugnyar, T. (2010): Do ravens show consolation? Responses to Distressed Others. Plos One, 5(5), e10605 Initiates file download 
  • Fraser, O. N., Bugnyar, T. (2010): The quality of social relationships in ravens. Animal Behaviour, 79, 927-933  Initiates file download
  • Hemetsberger, J., Scheiber, I.B.R., Weiß, B.M., Frigerio, D., Kotrschal, K. (2010): Influences of socially involved hand-raising on life history and stress responses in greylag geese. Interaction studies, 11 (3), 380-395
  • Hirschenhauser, K., Weiß, B.M., Haberl, W., Möstl, E., Kotrschal, K. 2010. Female androgen patterns and within-pair testosterone compatibility in domestic geese (Anser domesticus). Gen. Comp. Endocrinol., 165, 195-203 Initiates file download 
  • Kotrschal, K., Scheiber, I.B.R., Hirschenhauser, K. (2010): Individual performance in complex social systems. In: Animal Behaviour: Evolution & Mechanism (ed. by P. Kappeler). Heidelberg: Springer Verlag; pp. 121-148
  • Kralj-Fiser, S., Weiß, B.M., Kotrschal, K (2010): Behavioural and physiological correlates of personality in greylag geese (Anser anser). Journal of Ethology
  • Kralj-Fiser, S., Scheiber, I.B.R., Kotrschal, K., Weiß, B.M., Wascher, C.A.F. (2010): Glucocorticoids enhance and suppress heart rate and behaviour in time dependent manner in greylag geese (Anser anser). Physiology and Behaviour; 100, 394-400
  • Loretto, M.-C., Schloegl, C., Bugnyar, T. (2010): Northern bald ibises follow others' gaze into distant space but not behind barriers. Biology letters; 6, 14-17
  • Scheiber, I.B.R. (2010): Sozialverhalten im Lichte der Evolutionstheorie und Verhaltensbiologie. In: Evolution: Natur - Mensch - Gesellschaft. Wien: Peter Lang Verlag
  • Wascher, C.A.F., Fraser, O.N., Kotrschal, K. (2010): Heart Rate during Conflicts Predicts Post-Conflict Stress-Related Behavior in Greylag Geese. PLoS ONE 5(12): e15751. doi:10.1371/journal.pone.0015751 Initiates file download 
  • Wedl, M., Schöberl, I., Bauer, B., Day, J., Kotrschal, K. (2010): Relational factors affecting dog social attraction to human partners. Interaction studies; 11(3), 482-503
  • Weiß, B. M., Kotrschal, K., Möstl, E., Hirschenhauser, K. (2010): Social and life-history correlates of hormonal partner-compatibility in Greylag geese (Anser anser). Behavioral Ecology, 21, 138-143 (journal link)
  • Weiß, B.M., Kehmeier, S., Schloegl, C. (2010): Transitive inference in free-living greylag geese (Anser anser). Animal Behaviour, 79, 1277-1283


  • Bertin, A., Hirschenhauser, K., Kotrschal, K. (2009): Trans-generational influence of human disturbances in Japanese quail: egg quality influences male social and sexual behavior. Ethology; 115, 879-887  Initiates file download
  • Bertin, A., Richard-Yris, M.A., Houdelier, C., Richard, S., Lumineau, S., Kotrschal, K., Möstl, E. (2009): Divergent selection for inherent fearfulness leads to divergent yolk steroid levels in quails. Behaviour; 146(6), 757-770 Initiates file download 
  • Day, J.E.L., Kergoat, S., Kotrschal, K. (2009): Do pets influence the quantity and choice of food offered to them by their owners: lessons from other animals and the pre-verbal human infant? CAB Reviews: Perspectives in Agriculture, Veterinary Science, Nutrition and Natural Resources, 4, 42, 1-12 Initiates file download 
  • Kotrschal, K. (2009): Die evolutionäre Theorie der Mensch-Tier-Beziehung. In: Gefährten - Konkurrenten - Verwandte. Die Mensch-Tier-Beziehung im wissenschaftlichen Diskurs (ed. by. C. Otterstedt & M. Rosenberger). Göttingen: Vandenhoeck & Ruprecht Initiates file download 
  • Kotrschal, K., Schöberl, I., Bauer, B., Thibeaut, A.-M., Wedl, M. (2009): Dyadic relationships and operational performance of male and female owners and their male dogs. Behavioural Processes, 81, 383-391
  • Scheiber, I.B.R., Kotrschal, K., Weiß, B.M. (2009): Serial agonistic attacks by greylag goose families (Anser anser) against the same target. Animal Behaviour, 77, 1211-1216  Initiates file download
  • Scheiber, I.B.R., Kotrschal, K., Weiß, B.M. (2009): Benefits of family reunions: Social support in secondary greylag goose families. Hormones & Behavior 55, 133 – 138
  • Schloegl, C., Bugnyar, T., Aust, U. (2009): Exclusion performances in non-human animals: from chimpanzees to pigeons and back again. In: Rational animals, irrational humans (ed. by A. Blaisdell, L. Huber, S. Watanabe, Young, A.,  and Y. Yamazaki); Tokyo: Keio University Press; pp. 217-234
  • Schloegl, C., Dierks, A., Gajdon, G.K., Huber, L., Kotrschal, K., Bugnyar, T. (2009): What you see is what you get? Exclusion performances in ravens and keas; PLoS One, 4(8), e6368 Initiates file download 
  • Wascher, C.A.F., Scheiber, I.B.R., Weiß, B.M., Kotrschal, K. (2009): Heart rate responses to agonistic interactions in greylag geese, Anser anser. Animal Behaviour, 77, 955-961 Initiates file download 
  • Wedl, M., Kotrschal, K. (2009): Social and Individual Components of Animal Contact in Preschool Children; Antrozoös; 22(4), 383-396
  • Wedl, M., Bauer, B., Dittami, J., Schöberl, I., Kotrschal, K. (2009): Effects of Personality and Sex on Behavioral Patterns in Human-Dog Dyads. Journal of Veterinary Behavior: Clinical Applications and Research, 4 (2), 88-89 (published abstract) Initiates file download 


  • Bertin, A., Richard-Yris, M.-A., Houdelier, C., Lumineau, S., Möstl, E., Kuchar, A., Hirschenhauser, K., Kotrschal, K. (2008): Habituation to humans affects yolk steroid levels and offspring phenotype in quail. Hormones and Behavior, 54, 396-402 Initiates file download 
  • Bokony V., Garamszegi L.Z., Hirschenhauser K., Liker A. (2008): Testosterone and melanin-based black plumage coloration: a comparative study. Behavioural. Ecology and Sociobiology, 62, 1229-1238  Initiates file download 
  • Bugnyar, T. (2008): Rooks Team up to Solve a Problem. Current Biology, 18(12), R530-R531  Initiates file download
  • Garamszegi L.Z., Hirschenhauser K., Bokony V., Eens M., Hurtrez-Bousses S., Møller A.P., Oliveira R.F., Wingfield J.C. (2008): Latitudinal distribution, migration and testosterone levels in birds. American Naturalist, 172, 4, 533-546  Initiates file download
  • Hirschenhauser K., Canário A.V.M., Ros, A.F.H., Taborsky M., Oliveira R.F. (2008): Social context may affect urinary excretion of 11-ketotestosterone in African cichlids. Behaviour, 145, 1367-1388 Initiates file download 
  • Hirschenhauser K., Wittek M., Johnston P., Möstl E. (2008): Social context rather than behavioral output or winning modulates postconflict testosterone responses in Japanese quail (Coturnix japonica). Physiology and Behaviour, 95, 457-463 Initiates file download 
  • Kotrschal, K. (2008): Geist als Anpassung? In: Naturalisierung des Geistes? Beiträge zur Gegenwärtigen Debatte um den Geist. (K. Appel,  H.P. Weber, R. Langthaler & S. Müller eds.), pp. 84-92, Würzburg: Königshausen & Neumann
  • Kotrschal, K. (2008): Angepasst woran? Vom Mängel- zum Konfliktwesen. Pp. 55-65. In: Der Mensch – Ein Mängelwesen? Endlichkeit-Kompensation-.Entwicklungen. (H. Schmidinger & C. Sedmak, eds.) Darmstadt: WBG
  • Kotrschal, K. (2008): Menschen und Tiere: Eine evolutionäre Schicksalsgemeinschaft. In: Was ist der Mensch? Montagsakad. Bd. 4, pp. 125-135. Leykam Graz (
  • Range, F., Bugnyar, T., Kotrschal, K. (2008): The performance of ravens on simple discrimination tasks: a preliminary study. Acta Ethologica, 11, 34-41 Initiates file download 
  • Scheiber, I. B. R., Weiß, B. M., Hirschenhauser, K., Wascher, C. A. F., Nedelcu, I. T., Kotrschal, K. (2008): Does 'Relationship Intelligence' Make Big Brains in Birds? The Open Biology Journal, 1, 6-8  Initiates file download  
  • Scheid, C., Bugnyar, T. (2008): Short-term observational spatial memory in Jackdaws (Corvus monedula) and Ravens (Corvus corax). Animal Cognition, 11 (4), 691-698  Initiates file download 
  • Scheid, C., Schmidt, J. Noë, R. (2008): Distinct patterns of food offering and co-feeding in rooks. Animal Behaviour, 76, 1701-1707 Initiates file download 
  • Schloegl, C., Kotrschal, K., Bugnyar, T. (2008): Modifying the object-choice task: Is the way you look important for ravens? Behavioural Processes, 77(1), 61-65
  • Schloegl, C., Kotrschal, K., Bugnyar, T. (2008): Do common ravens (Corvus corax) rely on human or conspecific gaze cues to detect hidden food? Animal Cognition, 11, 231-241 Initiates file download 
  • Schloegl, C., Schmidt, J., Scheid, C., Kotrschal, K., Bugnyar, T. (2008): Gaze following in non-human animals: The corvid example. in: Animal Behaviour: New Research (ed. E. A. Weber & L. H. Krause), New York: Nova Science Publishers; pp. 73-92
  • Scholl, S., Grall, G., Petzl, V., Röthler, M., Slotta- Bachmayr, L., Kotrschal, K. (2008): Behavioural effects of goats on disabled persons. Therap. Comm. 2008
  • Schwab, C., Bugnyar, T., Schloegl, C., Kotrschal, K. (2008): Enhanced social learning between siblings in common ravens, Corvus corax. Animal Behaviour, 75, 501-508 Initiates file download 
  • Schwab, C., Bugnyar, T., Kotrschal, K. (2008): Preferential learning from non-affiliated individuals in jackdaws (Corvus monedula). Behavioural Processes, 79(3), 148-155
  • Scott A.P., Hirschenhauser K., Bender N., Oliveira R.F., Earley R.L., Sebire M., Ellis T., Pavlidis M., Hubbard, P.M., Huertas, M., Canário A.V.M. (2008): Non-invasive measurement of steroids in fish-holding water: important considerations when applying the procedure to behaviour studies. Behaviour, 145, 1307-1328 Initiates file download 
  • Stöwe, M., Bugnyar, T., Schloegl, C., Kotrschal, K., Heinrich, B. (2008): Corticosterone excretion patterns and affiliative behaviour over development in ravens (Corvus corax). Hormones & Behavior, 53, 208-216
  • Wascher, C. A. F., Arnold, W., Kotrschal, K. (2008): Heart rate modulation by social contexts in Greylag geese (Anser anser). Journal of Comparative Psychology, 122 (1), 100-107 Initiates file download 
  • Wascher, C.A.F., Scheiber, I.B.R., Kotrschal, K. (2008): Heart rate modulation in bystanding geese watching social and non-social events. Proceedings of the Royal Society of London, Series B, 275, 1653-1659 Initiates file download 
  • Weiß, B. M., Kotrschal, K., Frigerio, D., Hemetsberger, J., Scheiber, I. B. R. (2008): Birds of a feather stay together: extended family bonds, clan structures and social support in greylag geese (Anser anser). in: Family relations: behavioral, psychological and sociological aspects (ed. Columbus, F.), New York: Nova Science Publishers; 69-88
  • Weiß B.M., Poggemann K., Olek K., Foerster, K., Hirschenhauser K. (2008): Isolation and characterization of microsatellite marker loci in the greylag goose (Anser anser). Molecular Ecology Resources, 8, 1411-1413


  • Bugnyar, T. (2007): An integrative approach to the study of 'theory-of-mind'-like abilities in ravens. The Japanese Journal of Animal Psychology, 57(1), 15-27 Initiates file download 
  • Bugnyar, T., Schwab, C., Schloegl, C., Kotrschal, K., Heinrich, B. (2007): Ravens judge competitors through experience with play caching; Current Biology, 17, 1804-1808 Initiates file download 
  • Bugnyar, T., Stöwe, M., Heinrich, B. (2007): The ontogeny of caching in ravens, Corvus corax. Animal Behaviour, 74, 757-767 Initiates file download 
  • Kotrschal, K. (2007): Geistige Umweltverschmutzung. In: Macht um Welt. Pp. 27-37. BAWAG P.S.K.-Ed. Lit. ISBN 978-3-8000-7332-0. Ueberreuter (2007)
  • Kotrschal, K., Schloegl, C., Bugnyar, T. (2007): Instinkt oder Verstand? Lektionen von Rabenvögeln und Gänsen. Biologie in unserer Zeit, 37 (6), 366-374
  • Kralj-Fišer, S., Scheiber, I. B. R., Blejec, A., Moestl, E., Kotrschal, K. (2007): Individualities in a flock of free-roaming greylag geese: Behavioral and physiological consistency over time and across situations. Hormones and Behavior, 51, 239-248
  • Scheid, C., Range, F., Bugnyar, T. (2007): When, what and whom to watch? Quantifying attention in ravens and jackdaws. Journal of Comparative Psychology, 121, 380-386  Initiates file download 
  • Schloegl, C., Kotrschal, K., Bugnyar, T. (2007): Gaze following in Common ravens Corvus corax: ontogeny and habituation. Animal Behaviour, 74, 769-778  Initiates file download
  • Stöwe, M., Kotrschal, K. (2007): Behavioural phenotypes may determine whether social context facilitates or delays novel object exploration in ravens. Journal of Ornithology, 148 (Suppl 2), S179-184 Initiates file download 
  • Zoufal, K., Fritz, J., Bichler, M., Kirbauer, M., Markut, T., Meran, I., Wolf, A., Kotrschal, K. (2007) Feeding ecology of the Northern Bald Ibis in different habitat types: an experimental field study with handraised individuals. Report of the 2nd IAGNBI Meeting 2nd IAGNBI Meeting, Vejer 2006, C. Böhm, C. Bowden (Eds.), pp. 77-84.


  • Bugnyar, T., Heinrich, B. (2006). Pilfering ravens, Corvus corax, adjust their behaviour to social context and identity of competitors. Animal Cognition, 9, 369-376.  Initiates file download 
  • Hirschenhauser, K., Oliveira, R.F. (2006). Social modulation of androgens in male vertebrates: Meta-analyses of challenge hypothesis. Animal Behaviour 71, 265-277. Initiates file download 
  • Kotrschal, K., Hemetsberger, J., Weiß, B.M. (2006). Homosociality in male greylag geese (Anser anser): Making the best of a bad situation. In: Homosexual Behaviour in Animals: An Evolutionary Perspective (ed. P. Vasey and V. Sommer). Cambridge: Cambridge University Press; 45-76  Initiates file download
  • Range, F., Bugnyar, T., Schloegl, C., Schwab, C., Kotrschal, K. (2006). Individual learning ability and coping styles in ravens (Corvus corax). Behavioural Processes, 73(1), 100-106 Initiates file download 
  • Sorato E., Kotrschal, K. (2006). Hormonal and behavioural symmetries between the sexes in the Northern bald ibis. Journal of Comparative Endocrinology, 146, 265-274.  Initiates file download
  • Stöwe, M., Bugnyar, T., Loretto, M.-C., Schloegl, C., Range, F., Kotrschal, K. (2006). Novel-object exploration in ravens (Corvus corax): Effects of social relationships. Behavioural Processes, 73(1), 68-75 Initiates file download 
  • Stöwe, M., Bugnyar, T., Heinrich, B. and Kotrschal, K. (2006). Effects of Group Size on Approach to Novel Objects in Ravens (Corvus corax). Ethology, 112, 1079-1088 Initiates file download 


  • Bugnyar, T., Heinrich, B. (2005). Food-storing ravens differentiate between knowledgeable and ignorant competitors. Proceedings of the Royal Society of London, Series B 272, 1641-1646. Initiates file download 
  • Daisley, J.N., Bromundt, V., Möstl, E., Kotrschal, K. (2005). Enhanced yolk testosterone influences behavioural phenotype independent of sex in Japanese quail (Coturnix coturnix Japonica). Hormones and Behavior 47: 185-194.  Initiates file download
  • Heinrich, B., Bugnyar, T. (2005). Testing problem solving in ravens: String-pulling to reach food. Ethology 111, 962-976.  Initiates file download
  • Hemetsberger, J. (2005). Frühe Brutnachweise bei Steinlerche (Ammomanes isabellina), Saharasteinschmätzer (Oenanthe leucopyga), Akaziendrossling (Turdoides fulvus) und Hausammer (Emberiza striolata) in der zentralen Sahara Algeriens. Vogelwarte 43: 271-272. Initiates file download 
  • Hirschenhauser, K., Frigerio, D. (2005). Hidden patterns of male sex hormones and behavior vary with life history. In (Anolli, L., Duncan, S., Magnusson, M. and Riva, G., Eds.) The Hidden Structure Of Social Interaction, Chapter 5, pp. 81-96. Amsterdam: IOS-Press. Initiates file download 
  • Hirschenhauser, K., Kotrschal, K., Möstl, E. (2005). A synthesis of measuring steroid metabolites in goose feces. Annals of the New York Academy of Sciences 1046: 138-153. Initiates file download 
  • Scheiber, I. B. R., Weiß B. M., Frigerio D., Kotrschal K. (2005a). Active and passive social support in families of Greylag geese (Anser anser). Behaviour, 142, 1535-1557. Initiates file download 
  • Scheiber, I.B.R., Kralj S., Kotrschal K. (2005b). Sampling effort/frequency necessary to infer individual acute stress responses from fecal analysis in Greylag geese (Anser anser). Annals of the New York Academy of Sciences 1046, 154-167.  Initiates file download
  • Weiß, B.M., Möstl, E., Hirschenhauser, K. (2005). Within-pair testosterone compatibility as a currency for pairbond quality in Greylag geese? Hormones and Behavior 48, 133. Initiates file download 


  • Bugnyar, T., Kotrschal, K. (2004). Leading a conspecific away from food in ravens, (Corvus corax)? Animal Cognition 7, 69-76. Initiates file download
  • Bugnyar, T., Stöwe, M., Heinrich, B. (2004). Ravens, Corvus corax, follow gaze direction of humans around obstacles. Proceedings Royal Society London Series B 271, 1331-1336. Initiates file download 
  • Frigerio, D., Dittami, J., Möstl, E., Kotrschal, K. (2004a). Excreted corticosterone metabolites co-vary with ambient temperature and air pressure in male Greylag geese (Anser anser). General and Comparative Endocrinology 137, 29-36. Initiates file download 
  • Frigerio, D., Hirschenhauser, K., Möstl, E., Dittami, J., Kotrschal, K. (2004b). Experimentally elevated testosterone increases status signalling in male Greylag geese (Anser anser). Acta Ethologica 7, 9-18.  Initiates file download   
  • Goymann, W., Moore, I. T., Scheuerlein, A., Hirschenhauser, K., Grafen, A., Wingfield, J. C. (2004). Testosterone in tropical birds: effects of environmental and social factors. American Naturalist. 164, 327-334. Initiates file download 
  • Hirschenhauser, K., Taborsky, M., Oliveira, T., Canario, A.V.M., Oliveira, R.F. (2004). A test for the ‘challenge hypothesis’ in cichlid fish: simulated partner and territory intruder experiments. Animal Behaviour 68, 741-750. Initiates file download 
  • Hirschenhauser, K., Oliveira, R. F. (2004). Androgen responsiveness to the social environment: a comparative approach. Hormones and Behavior 46, 131-132.  Initiates file download 
  • Hemetsberger, J., Petutschnig, B. (2004). Zur Avifauna Jordaniens. Denisia 14: 421-436. Initiates file download 
  • Kotrschal, K., Bromundt, V., Föger, B. (2004). Faktor Hund - Hund. Eine sozio-ökonomische Bestandsaufnahme der Hundehaltung in Österreich. Czernin Verlag, Wien.
  • Scheiber, I.B.R. (2004). Social support as a key factor for individual success in Greylag geese (Anser anser)? Hormones and Behavior 46, 121.  Initiates file download
  • Weiß, B.M., Kotrschal, K. (2004). Effects of passive social support in juvenile Greylag geese (Anser anser): A study from fledging to adulthood. Ethology 110: 429-444. Initiates file download 


  • Frigerio, D., Weiss, B., Dittami, J., Kotrschal, K. (2003). Social allies modulate corticosterone excreation and increase success in agonistic interactions in juvenile hand-raised geese (Anser anser). Canadien Journal of Zoology ; 81:1746-1754  Initiates file download
  • Hackl, R., Bromundt, V., Daisley, J., Kotrschal, K., Möstl E. (2003) Distribution and origin of steroid hormones in the yolk of Japanese quail eggs (Coturnix coturnix Japonica). J. Physiol. B. 173, 327-331. Initiates file download 
  • Hirschenhauser, K., Winkler, H., Oliveira, R.F. (2003) Comparative analyses of male androgen responsiveness to social environment in birds: The effects of mating systems ant paternal incubation. Hormones and Behavior 43, 508-519. Initiates file download 
  • Kotrschal, K. (2003). Das Ende der Beliebigkeit - zum evolutionär-biologischen Fundament des Menschen. Österreichische Zeitschrift für Soziologie, 2003/2, 21-43 Initiates file download 
  • Kotrschal, K. (2003). Im Egoismus vereint? Zweite, überarbeitete Auflage. Fürth: Filander Verlag Fürth, 250pp, ISBN 3-930831-44-9. Initiates file download 
  • Kotrschal, K., Ortbauer, B. (2003). Behavioral effects of the presence of a dog in a classroom. Anthrozoös 16 (2), 147-159. Initiates file download 


  • Bugnyar, T., Kotrschal, K. (2002) Observational learning and the raiding of food caches in ravens (Corvus corax): Is it "tactical" deception? Animal Behaviour 64, 185-195. Initiates file download 
  • Bugnyar, T., Kotrschal, K. (2002). Scrounging tactics in free-ranging ravens, (Corvus corax). Ethology 108, 993-1009. Initiates file download 
  • Fritz, J., K. Kotrschal, K. (2002) On avian imitation: cognitive and ethological perspectives. In: Imitation in Animals and artifacts (Ed. by K. Dautenhahn and C. Nehaniv), MIT Press, Cambridge. S 133-157.
  • Hirschenhauser, K., Frigerio, D., Grammer, K., Magnusson, M.S. (2002) Monthly patterns of testosterone and behaviour in prospective fathers. Hormones and Behavior 42, 172-181. Initiates file download 
  • Kijne, M., Kotrschal, K. (2002) Neophobia affects choice of food-item size in group-foraging common ravens(Corvus corax). Acta Ethologica 5:13-18  Initiates file download
  • Kotrschal, K. (2002) Eine integrative, systemorientierte Sicht sozialer Mechanismen. In: Soziale Mechanismen. (U. Gansloßer ed.), Fürth: Filander Verlag
  • Pfeffer, K., Fritz, J., Kotrschal, K. (2002) Hormonal correlates of being an innovative greylag goose. Animal Behaviour 63, 687-695. Initiates file download 
  • Tintner, A., Kotrschal K. (2002) Early social influence on nestling development in Waldrapp ibises (Geronticus eremita). Zoo Biology 21, 467-480 Initiates file download 


  • Bugnyar, T., Kotrschal, K. (2001) Movement coordination and signalling in ravens (Corvus corax): an experimental field study. Acta Ethologica, 3, 101-109 Initiates file download 
  • Bugnyar, T., M. Kijne, Kotrschal K. (2001) Food calling in ravens: are "yells" referential signals? Animal Behaviour 61, 949-958. Initiates file download 
  • Frigerio, D., Möstl, E., Kotrschal, K. (2001) Excreted metabolites of gonadal steroid hormones and corticosterone in greylag geese (Anser anser) from hatching to fledging. General and Comparative Endocrinology 124, 246-255  Initiates file download
  • Frigerio, D., Weiss, B., Kotrschal, K. (2001) Spatial proximity among adult siblings in greylag geese (Anser anser): evidence for female bonding? Acta Ethologica, 3, 121-125 Initiates file download 
  • Fritz, J. (2001) Der intelligente Kumpan am Beispiel der Graugänse. In: Konzepte der Verhaltensforschung. Konrad Lorenz und die Folgen (ed. by K. Kotrschal, G. Müller & H. Winkler). Fürth: Filander Verlag
  • Hemetsberger, J. (2001) Die Entwicklung der Grünauer Graugansschar seit 1973. In: Konzepte der Verhaltensforschung. Konrad Lorenz und die Folgen (ed. by K. Kotrschal, G. Müller & H. Winkler). Fürth: Filander Verlag
  • Kotrschal, K. (2001) Das Rousseausche Vorurteil vom "edlen Wilden": Die Wurzeln der Ethik von Konrad Lorenz. In: Konzepte der Verhaltensforschung. Konrad Lorenz und die Folgen (ed. by K. Kotrschal, G. Müller & H. Winkler). Fürth: Filander Verlag. 109-117
  • Kotrschal, K. (2001) Der Beitrag von Österreichern zur Wissenschaft der Biologie vom Verhalten und zum evolutionären Bild des Menschen. In: K. Acham (ed.): Cognitio Humana. Zur Geistesgeschichte Österreichs.
  • Kotrschal, K., Bugnyar T., Stöwe, M. (2001) Kognition und Neophobie bei Raben. Charadrius 37, 127-134.
  • Möstl, E., Spendier H., Kotrschal K. (2001) Concentration of immunoreactive progesterone and androgens in the yolk of hen´s eggs (Gallus domesticus). Wiener Tierärztliche Monatsschrift 88, 62-65.
  • Weiß, B. M., Kotrschal, K. (2001). Social support in Greylag geese (Anser anser) from fledging to adulthood. In: Adv. Ethol. 36 (ed. Apfelbach, R.  Fendt, M., Krämer, S. and Siemers, B.M.), Berlin: Blackwell Sciences, 288

older publications

  • Fritz, J., Bisenberger, A., Kotrschal, K. (2000) Stimulus enhancement in greylag geese: socially mediated learning of an operant task. Animal Behaviour, 59, 1119-1125. Initiates file download 
  • Hemetsberger, J. (2000): Singvogel-Monitoring am Almsee, Oberösterreich. Erste Ergebnisse des Jahres 1999. Vogelkundliche Nachrichten Oberösterreichs - Naturschutz aktuell, 8 (1), 19-27  Initiates file download
  • Hirschenhauser, K., Möstl, E., Wallner, B., Dittami, J., Kotrschal, K. (2000) Endocrine and Behavioural Responses of Male Greylag Geese (Anser anser) to Pairbond Challenges during the Reproductive Season. Ethology, 106, 63-77  Initiates file download
  • Hirschenhauser, K., Möstl, E., Péczely, Wallner, B., Dittami, J., Kotrschal, K. (2000) Seasonal Relationships between Plasma and Fecal Testosterone in Response to GnRH in Domestic Ganders. General and Comparative Endocrinology, 118, 262-272 Initiates file download 
  • Kotrschal, K., Dittami, J., Hirschenhauser, K., Möstl, E., Péczely (2000) Effects of physiological and social challenges on fecal testosterone and corticosterone in male domestic geese (Anser domesticus). Acta Ethologica, 2, 115-122  Initiates file download
  • Fritz, J., Kotrschal, K. (1999) Social learning in common ravens, Corvus corax. Animal Behaviour, 57, 785-793.  Initiates file download 
  • Hirschenhauser, K., Möstl, E., Kotrschal, K. Within-pair testosterone covariation and reproductive output in Greylag Geese Anser anser. Ibis, 141, 577-586   Initiates file download 
  • Hirschenhauser, K., Möstl, E., Kotrschal, K. (1999) Seasonal Patterns of Sex Steroids Determined from Feces in Different Social Categories of Greylag Geese (Anser anser). General and Comparative Endocrinology, 114, 67-79 Initiates file download 
  • Hirschenhauser, K., Möstl, E., Kotrschal, K. (1999) Faecal analysis of androgens and its relevance for the study of social behaviour and reproductive success in Greylag geese Anser anser. Advances in Ethology, 34, 206
  • Kotrschal, K., Hirschenhauser, K., Möstl, E., Peczely, P. (1999) Testosterone and corticosterone determination from feces in geese yields biologically relevant results. Advances in Ethology, 34, 207
  • Kotrschal, K. (1999): Taste(s) and olfaction(s) in fish: a review of specialized sub-systems and central integration. Pflügers Archiv: European Journal of Physiology, 439 (7), r178-r180
  • Kotrschal, K. (1998): Sensory Systems. In: Intertidal Fishes (ed. by M. Horn, K. Martin & M. Chotkowski), San Diego: Academic Press; 126-142
  • Kotrschal, K., Hirschenhauser, K., Möstl, E. (1998) The relationship between social stress and dominance is seasonal in greylag geese. Animal Behaviour, 55, 171-176.  Initiates file download
  • Kotrschal, K., Van Staaden, M., J., Huber, R. (1998) Fish brains: evolution and environmental relationships. Reviews in Fish Biology and Fisheries, 8, 373-408.  Initiates file download 
  • Weiß, B., Kotrschal, K. (1997). Humans as social allies: interspecific social support in greylag geese (Anser anser). In: Adv. Ethol. 32 (ed. M. Taborsky and B. Taborsky); Berlin: Blackwell Sciences; 150
  • Hirschenhauser, K., Dittami, J., Möstl, E., Wallner, B., Kotrschal, K. (1997). Testosterone and behaviour during different phases of reproduction in greylag geese (Anser anser). In: Adv. Ethol. 32 (ed. M. Taborsky and B. Taborsky); Berlin: Blackwell Sciences; 61
  • Frigerio D., Hirschenhauser K., Wallner B., Kotrschal K. (1997). Experimentally increased testosterone affects behaviour and corticosterone in male greylag geese. In: Adv. Ethol. 32 (ed. M. Taborsky and B. Taborsky); Berlin: Blackwell Sciences; 59
  • Kotrschal, K. (1997): Die Konrad Lorenz Forschungsstelle in Grünau. Vogelkundliche Nachrichten Oberösterreichs - Naturschutz aktuell, 4, 31-33   Initiates file download
  • Kotrschal, K., Finger, T., E. (1996). Secondary Connections of the Dorsal and Ventral Facial Lobes in a Teleost Fish, the Rockling (Ciliata mustela). Journal of Comparative Neurology 370, 415-426  Initiates file download  
  • Kotrschal, K., Krautgartner, W.D., Hansen, A. (1997): Ontogeny of the solitary chemosensory cells in the zebrafish, Danio rerio. Chemical senses, 22 (2), 111-118  Initiates file download
  • Kotrschal, K. (1996): Solitary chemosensory cells: why do primary aquatic vertebrates need another taste system? Trends in Ecology & Evolution, 11(3), 110-114   Initiates file download
  • Kotrschal, K., Finger, T. (1996): Secondary connections of the dorsal and ventral facial lobes in a teleost fish, the rockling (Ciliata motella). Journal of Comparative Neurology, 370, 415-426
  • Kotrschal, K., Peters, R.C., Doving, K.B. (1996): Chemosensory and tactile nerve responses from the anterior dorsal fin of a rockling, Gaidropsarus vulgaris (Gadidae, Teleostei). Primary Sensory Neuron, 1, 297-308
  • Kotrschal, K. (1995). Ecomorphology of solitary chemosensory cell systems in fish: a review. Environmental Biology of Fishes, 44, 143-155 Initiates file download 
  • Kotrschal, K., Essler, H. (1995): Goals and approaches in the analysis of locomotion in fish, with a focus on laboratory studies. Reviews in Fisheries Science, 3, 171-200
  • Kotrschal, K., Hemetsberger, J. (1995). Social constraints in feeding and individual specialization for transportable food in greylag geese (Anser anser). Ökologie der Vögel, 17, 157-163 Initiates file download 
  • Essler, H., Kotrschal, K. (1994): High resolution analysis of swim path patterns of intact and olfaction-deprived minnows (Phoxinus phoxinus) stimulated with food and potential predator odour. Journal of Fish Biology, 45, 555-567
  • Kotrschal, K., Hemetsberger, J., Dittami, J. (1993). Food exploitation by a winter flock of greylag geese: behavioral dynamics, competition and social status. Behavioural Ecology and Sociobiology, 33, 289-295 Initiates file download 
  • Kotrschal, K., Peters, R., Atema J. (1993): Sampling and behavioral evidence for mucus detection in a unique chemosensory organ: the anterior dorsal fin in rocklings (Ciliata mustela: Gadidae: Teleostei). Zoologische Jahrbücher Abteilung fuer Allgemeine Zoologie und Physiologie der Tiere, 97, 47-67 
  • Kotrschal, K., Whitear, M., Finger, T., E. (1993) Spinal and Facial Innervation of the Skin in the Gadid Fish Ciliata mustela (Teleostei). Journal of Comparative Neurology, 331, 407-417  Initiates file download
  • Kotrschal, K., Hemetsberger, J., Dittami, J.(1992). Vigilance in a flock of semi-tame Greylag Geese Anser anser in response to approaching eagles Haliaeetus albicilla and Aquila chrysaetos. Wildfowl, 43, 215-219 Initiates file download 
  • Mausz, B., Dittami, J., Kotrschal, K. (1992): Triumphgeschrei und Aggression bei der Graugans (Anser anser). Ökologie der Vögel, 14, 165-172  Initiates file download 
  • Kotrschal, K. (1992): Quantitative scanning electron microscopy of solitary chemoreceptor cells in cyprinids and other teleosts. Environmental Biology of Fishes, 35, 273-282
  • Kotrschal, K., Palzenberger, M. (1992): Neuroecology of cyprinids: comparative, quantitative histology reveals diverse brain patterns. Environmental Biology of Fishes, 33, 135-152   Initiates file download
  • Finger, T., E., Drake, S., K., Kotrschal, K., Womble, M., Dockstader, K., C. (1991). Postlarval Growth of the Peripheral Gustatory System in the Channel Catfish - Ictalurus punctatus. Journal of Comparative Neurology, 314, 55-66 Initiates file download 
  • Kotrschal, K. (1991). Solitary chemosensory cells - taste, common chemical sense or what. Reviews in Fish Biology and Fisheries, 1, 3-22.  Initiates file download
  • Peters, R., Kotrschal, K., Krautgartner, W.-D. (1991): Solitary chemoreceptor cells of Ciliata mustela (Gadidae, Teleostei) are tuned to mucoid stimuli. Chemical senses, 16, 31-42
  • Kotrschal, K. (1989) Trophic ecomorphology in eastern Pacific blennioid fishes - character transformation of oral jaws and associated change of their biological roles. Environmental Biology of Fishes, 24 (3), 199-218 Initiates file download 
  • Kotrschal, K., Whitear, M. (1988): Chemosensory anterior dorsal fin in rocklings (Gaidropsarus and Ciliata, teleostei, Gadidae): Somatropic representation of the ramus recurrens facialis as revealed by transganglionic transport of HRP. The Journal of Comparative Neurology, 268(1), 109-120  Initiates file download 
  • Kotrschal, K. (1988) Evolutionary patterns in tropical marine reef fish feeding. Journal of Zoological Systematics and Evolutionary Research, 26, 51-64  Initiates file download  
  • Lindquist, David G., Kotrschal, K. (1987) The Diets in Four Pacific Tube Blennies (Acanthemblemaria: Chaenopsidae): Lack of Ecological Divergence in Syntopic Species. Marine Ecology, 8 (4), 327-335  Initiates file download  
  • Kotrschal, K., Lindquist, D. G. (1986) The Feeding Apparatus in four Pacific Tube Blennies (Teleostei: Chaenopsidae): Lack of Ecomorphological Divergence in Syntopic Species. Marine Ecology, 7 (3): 241-253  Initiates file download
  • Goldschmid, A., Kotrschal, K. (1985) Morphological and functional adaptations in different feeding types of blennies (Perciformes; Teleostei). Fortschritte der Zoologie, 30, 241-244  Initiates file download

Konrad Lorenz Forschungsstelle
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4645 Grünau

T: +43 7616 8510
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